61 



ophthalmica magna of fishes. There is no orbitonasalis branch of the 

 carotis interna, and also no palatine branch. 



In mammals, the post-trigeminus portion of the primary lateral 

 wall of the trigemino-facialis chamber, already separated from the pre- 

 trigeminus portion in Lacerta, has become definitely related to the 

 auditoiy capsule, and hence forms no part of the lateral wall of the 

 persisting chamber (cavum epiptericum, Gafpp). The post-trigeminus 

 portion of the lateral wall of the latter chamber must accordingly 

 be of secondary origin, which is wholly in accord with the fact that 

 it is, in Echidna, (Gaupp, 1908), and hence probably also in other 

 vertebrates also, of relatively late development and first appears as a 

 membrane which later ossifies without previous chondrification. This 

 portion of the lateral wall of the cavum epiptericum of mammals is 

 accordingly in no way the homologue of the corresponding portion of 

 the wall of the trigemino-facialis chamber of fishes, this doubtless ac- 

 counting for the fact that the carotis externa, in mammals, does not tra- 

 verse the chamber, so far as I can find, as it does in lower vertebrates. 



Palais de Carnol^s, Menton, May 29 th. 1914. 



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