THE EOCENE: PERIOD. 301 
by his researches to form as to the outward appearance of 
Paleotherium magnum. Recent discoveries, however, have 
= = Fae My 
We as & ae With <= 
ea - : = 
fee ———— ae = ZEW" «gni7 i ES ax) 
—— 
Fig .229 —Outline of Padeotherium magnum, restored. Upper Eocene, Europe. 
fter Cuvier.) 
rendered it probable that this restoration is in some important 
respects inaccurate. Instead of being bulky, massive, and 
more or less resembling the living Tapirs in form, it would rather 
appear that Paleotherium magnum was in reality a slender, 
graceful, and long-necked animal, more closely resembling in 
general figure a Llama, or certain of the Antelopes. 
The singular genus Anchithertum forms a kind of transition 
between the Pa/gotheria and the true Horses (Zguide). The 
Horse (fig. 230, D) possesses but one fully-developed toe to 
each foot, this being terminated by a single broad hoof, and 
representing the mzddle toe—the ¢hird of the typical five- 
fingered or five-toed limb of Quadrupeds in general. In 
addition, however, to this fully-developed toe, each foot in 
the horse carries two rudimentary toes which are concealed 
beneath the skin, and are known as the “splint-bones.” 
These are respectively the second and fourth toes, in an 
aborted condition; and the fst and f/th toes are wholly 
wanting. In Azpparion (fig. 230, C), the foot is essentially 
like that of the modern Horses, except that the second and 
fourth toes no longer are mere “splint-bones,” hidden be- 
neath the skin; but have now little hoofs, and hang freely, 
but uselessly, by the side of the creat middle toe, not being 
sufficiently developed to reach the ground. In Anchitherium, 
again (fig. 230, B), the foot is three-toed, like that of Azpparion, 
but the two lateral toes (the second and fourth) are so far 
