ARCHER, ON SAPROLEGNIEX. 125 
sufficiently early to see the evolution of the zoospores, and 
thus to determine the genus. Still, combining two indica- 
tions furnished from other sources, presently to be mentioned, 
the evidence seems sufficiently strong to point to the genus 
Achlya as the proper location of this species. 
I have mentioned that this form is dicecious, but I had the 
good fortune to meet with the empty mother-cells only of the 
spermatozoids. Their structure and mode of development, 
however, agreed so completely with that part of the fructifi- 
cation in Achlya dioica, as figured by Pringsheim, that there 
is no need here to give a drawing.* A terminal portion of 
one of the tubular filaments of which the plant is com- 
posed was divided by transverse septa into several cavities, 
two or three times longer than broad; these cavities were 
densely filled by empty globular hyaline coats, which had 
evidently been evacuated by the contents. The only differ- 
ence from Pringsheim’s figure consisted in these special- 
mother-cells being somewhat smaller and more numerous. 
Now, whereas in Achlya dioica (Pringsh.) the spermatozoids 
are produced by unskinning from a special mother-cell, as 
are also the zoospores, so also I think we may feel justified 
in assuming from analogy, inasmuch as the spermatozoids in 
the present instance are formed by unskinning from a special 
mother-cell, that likewise here too are the zoospores. If so, 
this plant would fall under the genus Achlya. 
Another indication pointing to the genus Achlya is as 
follows : 
In this new species, not infrequently just under a terminal 
oogonium, the main filament gives off one or two or three 
lateral branches in a kind of proliferous manner, and these 
are usually of considerably less diameter than that of the 
supporting stem. These, at first sight, might be supposed to 
look not unlike what might be intended to become lateral 
male branchlets (fig. 5), sufficiently puzzling after one has 
previously found that the species is a diccious one. But 
when we notice that the oospores are here fully formed, and 
yet that this lateral branch still retains its contents and is not 
in contact with the oogonium, such a mistake is prevented. 
Such a form as that drawn in fig. 6 at once, however, ex- 
plains the former case. Here we see the ends of these 
become inflated, densely filled with contents, and shut off as 
oogonia. In these secondary oogonia I never noticed more 
than one oospore, although the first-formed oogonia might con- 
tain perhaps as many as eight or ten, though ordinarily fewer. 
Now, this proliferous manner of growth is the second cir- 
* ‘ Jahrbiicher fiir wiss. Bot.,’ Band ii, t. xxiii, fig. 2. 
