STRUCTURE OF THE MAMMALIAN GASTRIC GLANDS. 381 



It has seemed to me that there is an increase of indulino- 

 philous cells in the body of the gland during the first hours of 

 digestion^ but I have not yet given this matter enough atten- 

 tion to enable me to speak with certainty concerning it. 



The discovery of the different nature of the cells of the 

 neck of the gland affords a cytological basis for the division of 

 the gland into two regions, a neck and body. 



The neck of the gland^ as determined by the distribution of 

 the indulinophilous cells, varies in length in the different por- 

 tions of the stomach. It is shortest in the glands of the 

 greater curvature, where it forms about one third of the gland, 

 and longest in the lesser curvature, where it may comprise as 

 much as four-fifths of the entire gland. 



B. The Pyloric Glands of the Cat. 



These glands are, as Toldt^ pointed out, branched tubular 

 glands, consisting of a deep pit or duct lined by a continua- 

 tion of the surface epithelium, and, opening into this, branched 

 tubules of various lengths exhibiting a tortuous course in the 

 deeper layers of the mucosa. 



Disregarding the anatomical divisions of the gland, one may 

 divide it, on a basis of the nature of the cells, into three 

 regions : a funnel-shaped duct, lined by a continuation of the 

 surface epithelium (fig. 7, a) ; the tubular body of the gland, 

 lined by the true pyloric gland cells (fig. 7, c) ; and a short por- 

 tion connecting these, and lined by cells of an intermediate 

 type (fig. 7, b). 



The true pyloric gland cells are, except as regards shape, 

 identical with the chief cells of the neck of the fundus gland. 

 They present all the characteristic staining properties of the 

 latter, and, if one compares similar secretion phases, are of 

 the same structure. They contain a secretion which stains in- 

 tensely in Bordeaux R, in indulin and Mayer's muchaematein, 

 and, as R. Krause has already pointed out, gives a meta- 

 chromatic red stain in thionin. This secretion I regard, for 

 similar reasons, as of a mucous nature. 



' ' Sil.zuiigsber. d. k, Akad. d. VVissenscli., Wien,' 1881. 



