84- Mr. B. Clarke on Relative Position ; 



already noticed as occurring in Cornus and Marlea, and the fol- 

 lowing enumeration will show that it is one of the more fre- 

 quent variations of the position of the raphe. 



1. Pontedera lanceolata. 2. Aquifoliacese {Ilex). 3. Styra- 

 ceae [Halesia — ovules two). 4. Oleaceae (ovules two). 5. Mal- 

 pighia and other genera of Malpighiacese in which the funiculus 

 (representing the rajihe) is constantly lateral (PI. II. fig. 5). 

 6. A^zYrflWa as figured by Prof. Lindley. 7. Santalaceee. 8. Myo- 

 poraceae (ovule single or two). 9. Illecebracese. 10. Cheno- 

 podiacese. 11. Epacridete [Acrotriche) . 12. Caprifoliaceje. 

 13. Globuluria. 14. Dipsacese. 15. Valerianacese. 16. Hip- 

 puridese {Goniocarpus) . 17. HamamelideEe. 18. Bruniacese. 

 19. Schizandra, 



In Corrigiola the cotyledons are lateral, that is, neither next 

 the funiculus nor yet directly removed from it, and therefore if 

 the ovule were completely inverted the raphe woiild be lateral. 

 In Paronychia, however, the cotyledons are turned away from 

 the funiculus and the radicle in relation with it ; but as the raphe 

 averse, and the raphe lateral, occur in the same family, as in 

 Aucuba and Cornus, and equivalent characters also in Malpi- 

 ghiacese, this oflFei's an explanation of the variable relation of the 

 cotyledons to the funiculus in Illecebracese, and also tends to 

 show that the raphe averse the placenta and the cotyledons 

 averse it [i. e. next the dorsum of the cell) are characters of 

 equivalent value. 



3. Ovule pendulous with the ra]jhe next the placenta. This, as 

 is well known, is the ordinary position of the raphe in pendulous 

 anatropal ovules ; but although it is the more common, it is of 

 rare occurrence in the Heterocarpous families, as will be seen 

 from the Tables. 



4. Ovule horizontal with the raphe on the upper surface. Of 

 this position I have hitherto observed only three instances. 



1 . A species of Macleya in which the ovules are six, three on 

 each placenta, having the raphe constantly on the upper surface. 



2. Aracese. Where they are numerous, many of the ovules have 

 this character, but in others the raphe is lateral. 3. Fumaria 

 officinalis. The ovule, although not anatropal, has the equivalent 

 character of the foramen, being always directly below its attach- 

 ment to the wall of the ovary. To these perhaps should be 

 added Paris quadrifulia as the ovules are scarcely ascending, the 

 raphe being frequently on the upper surface but sometimes late- 

 ral, thus agreeing with Aracese; and also the ovules on the upper 

 portion of the placenta in Swietiana, where in the early stages 

 they are horizontal. 



5. Ovule horizontal with the raphe lateral. Of this no instance 

 has been observed where the ovule is single, except in Chenopo- 



