270 3Ir. Bentham's Review of the Order of Hydrophyllese. 



in tlie Lab'iativ, Echium in Borraginece, or Eutoca and Phacelia in Hydro- 

 pkyllecE, being present or absent in two species otherwise very closely allied. 

 In Hijdrophyllece, their form is very variable. In general their centre is entirely 

 blended with the corolla, and their broad dilated margins, embracing the basis 

 of the filaments, are alone visible ; but in the genus Hydrophyllum they appear 

 to be constantly linear, adnate along the back, but free at the upper extremity 

 and the margins. In Emmenanthe and some Eutocce, as also in Echium viil- 

 gare and several Cynoglossa, they are reduced to ten very small orbicular 

 squamae, placed quite at the base of the corolla ; and in Eutoca grandijiora 

 and parviftora, Phacelia Jimhriata, and some others, they disappear entirely, 

 a transverse nerve connecting the base of the stamina alone indicating their 

 usual position. 



The stamina in all Hydrophyllece are much alike, of equal size, and regularly 

 divergent; their only differences are in the unimportant character of length and 

 in the hairiness of the filaments. The anthers are always oblong or linear, 

 with parallel cells. 



The style, in several species, shows readily to the naked eye the real struc- 

 ture of those Labiatce, Borragitiece, &c., which are said to have a simple style, 

 with two subulate stigmata. The style is, on the contrary, in fact bifid, each 

 lobe bearing at the extremity a small stigma. The ovarium of Emmenanthe 

 is covered with a glandular jiubescence ; that of all other Hydrophyllece is 

 clothed with white erect rigid hairs. The style of Phacelia and Eutoca is 

 usually more deeply cleft than in the other genera, but the latter character 

 is very uncertain. 



The placentation of the ovarium is of great importance in the generic dis- 

 tribxition of Hydrophyllea;. In Hydrophyllum, Nemophila, and Ellisia the two 

 placentae are broad, fleshy, line the whole ovarium, adhere at the top and basis 

 only, being fi-ee from the parietes, and bear on their inner siu'face each of them 

 from two to sixteen ovulse placed in two vertical rows, one on each side of the 

 central line. In Eutoca, Phacelia, and Emmenanthe the placenta; are linear or 

 slightly dilated, and adhere more or less to the parietes along their central line, 

 bearing on their inner surface from two to fifty or sixty ovulse, arranged either 

 in two rows, or covering the whole surface without any apparent arrangement. 



As the fruit ripens, the broad placentae of the three first-named genera con- 



