320 CALIFORNIA ACADEMY OF SCIENCES. [Proc. 3D Ser. 



fertilized; in S. Greenii they remain separate until after 

 fertilization. 



3. The antipodal cells are three in number and in S. 

 simplex 2ire. very inconspicuous; after fertilization there is 

 a remarkable secondary growth in the antipodal cells, re- 

 sulting finally in a large cell-mass, containing in S . simplex 

 over 150 cells. 



4. The process of fertilization, so far as it was studied, 

 offered no anomalies. 



5. The development of the embryo in S. simplex agrees 

 closely with that of S. ramosum, studied by Hegelmaier. 

 There are regularly three primary transverse segments, of 

 which the terminal one gives rise to cotyledon, stem-apex, 

 and part of the root. The suspensor remains undeveloped, 

 and in this respect the embryo is like that of the Graminese. 

 The slight development of the suspensor is associated with 

 the complete investment of the young embryo by the endo- 

 sperm. 



6. The stem-apex is lateral in origin like that of most 

 Monocotyledons, but is not developed from the middle of the 

 three primary segments. 



7. The primary tissues of the embryo are very early 

 developed, especially in the terminal embryonal segment. 

 The plerome of the root is derived entirely from the terminal 

 segment; the initials for the other tissues of the root arise 

 from the middle segment. 



8. The development of the endosperm follows the usual 

 course, but is rather late in forming the first division-walls. 

 The enlarged antipodal cells doubtless function at first as 

 endosperm. 



9. The large endosperm-nuclei have often several nucle- 

 oli which are formed by a fragmentation of the original 

 nucleolus. 



10. Large crystalloids are abundantly developed in the 

 older endosperm-cells of S . simplex. 



11. The inner pericarp develops into sclerenchyma 

 with thick, deeply pitted walls. The development of the 

 " Samen-deckel " is the same as described by Hegelmaier 

 for S. ramosum. 



