FORT UNIOX OF CRAZY MOUNTAIN FIELD, MONT. 131 



On the other hand, the following characters show that the relation- 

 ship cannot be very close, if it exists at all: 



Rather than a single mesostyle developed as a fold of the ectoloph, the plagio- 

 menids have two independent median cusps on the outer shelf of the upper molars. 



The anterior incisors (in the lower jaw, at least) are not reduced in number and 

 none is markedly enlarged. 



The hypoconulid does not have the characteristic mixodectid displacement 

 toward the entoconid. 



The cheek teeth are all deeply furrowed and tend to proliferate cuspules. 



The inconclusive evidence of possible relationship of the plagio- 

 menids to the Dermoptera has not been significantly altered since 

 Matthew wrote (1918). The chain of evidence thus tending to link 

 the mixodectids with the Dermoptera is so weak at every point as 

 not to merit serious consideration at present. 



Genus EUDAEMONEMA Simpson 



Eudaemonema Simpson, 1935d, p. 231. 



Type. — Eudaemonema cuspidata Simpson. 



Distribution. — Middle Paleocene, Fort Union, Mont. 



Diagnosis. — Dental formula, 2.1.4.3. Median incisor enlarged. Canine 

 larger than lateral incisor or Pi. Pi_2 small, 1-rooted. P4 submolari- 

 form, with small paraconid, large, high metaconid, and basined, 

 tricuspid talonid. Lower molars as in Mixodedes, but trigonids more 

 elevated and all six cusps sharper and more distinct. Upper molars 

 with prominent hypocones, projecting strongly internally. 



EUDAEMONEMA CUSPIDATA Simpson 



Figures 25, 26 



Eudaemonema cuspidata Simpson, 1935d, p. 231. 



Type. — U.S.N.M. no. 9314, left lower jaw with C, P2-M3, and roots 

 or alveoli of all other teeth. Found by Dr. J. W. Gidley. 



Horizon and locality. — Gidley Quarry (referred specimen from 

 Silberling Quarry), Fort Union, Middle Paleocene horizon, Crazy 

 Mountain Field, Mont. 



Diagnosis. — Sole known species of genus. Dimensions in table 29. 



Remarks. — Alveoli in the type and roots in another specimen (no. 

 9317) show clearly that there was an enlarged median incisor with a 

 procumbent, laterally compressed root, flanked by one much smaller 

 lateral incisor. The canine root is oval, implanted nearly vertically^ 

 nearly as large as that of the median incisor, and much larger than 

 that of either of the adjacent teeth. The crown is low, with a simple^ 

 spatulate, recurved tip. Pi is not known, but its alveolus in three 

 different specimens suggests that it was constantly present, although 

 absent in all other known members of this family. Its root is single, 

 small, and circular in section. 



