488 PROF, MILNES MARSHALL AND W. B. SPENCER. 
the branches being few in number at the proximal end, and 
much more numerous distally. A short distance before its 
termination this nerve, like the ophthalmic branch of the fifth, 
is crossed at a considerable angle by the nerve JW (fig. 12). 
The connecting branch (WN. c’.) between the seventh and fifth 
nerves is well seen in figs. 11 and 12; it is now shorter and 
wider than at stage t (fig. 10), and contains very numerous 
ganglion cells along its whole length. 
The superficial portion of this nerve (vit @) is not shown in 
fig. 11, but is represented along its whole length in fig. 12 ; it 
is a stout nerve with a remarkably straight course ; it gives off 
no branches at all along the greater part of its length, but near 
its distal termination divides rather suddenly into a number of 
branches, which end in the integument of the maxillary process, 
the most anterior of the branches extending forwards almost as 
far as the hinder border of the olfactory pit. The relations of 
this nerve to the maxillary branch of the fifth are very curious ; 
the two nerves are very close together, the branch of the seventh 
lying, as at the earlier stage, immediately superficial to that of 
the fifth. These relations are well seen in the transverse section 
drawn in fig. 6. ‘This shows, as already noticed, the primary 
and secondary roots of the seventh, and also the proximal portion 
of the nerve vir d. This nerve is seen to be a direct continua- 
tion of the primary root; its inner, or deeper, portion is seen on 
both sides to become continuous with the maxillary branch of 
the fifth, (v 2), the junction of the two forming the connection 
between the fifth and seventh nerves already noticed. Beyond 
this point of union the nerve vit d is continued downwards, lying 
immediately superficial to the maxillary nerve (v4). The two 
nerves preserve this relation up to their terminal distribution, 
two of the ultimate branches being represented in figure 5 (v 4 
aud vit d@). This very remarkable branch of the seventh puzzled 
us greatly for a long time, and it was only after working out the 
whole history of its development up to stage q that we succeeded 
in determining its import. The nerve has already been described 
and figured by one of us,’ and named tentatively the palatine. 
This determination now proves to be erroneous; the palatine is a 
deep-lying nerve, whereas the nerve vir d retains its superficial 
position in the adult. 
This nerve (vt d) we have now identified as the buccal nerve, 
the proof of this determination, consisting in our having traced 
the nerve directly up to the adult. The buccal nerve has always 
hitherto been regarded as a branch of the fifth, and is described 
1 Marshall, loc. cit., pp. 86, 87, and Pls. V, fig. 15; and VI, figs. 28 
and 29. 
