558 DR. E. KLEIN. 
infiltrated with lymph-corpucles; this subepithelial layer is 
limited by a thin layer of elastic fibres running chiefly in a 
longitudinal direction, and it is this elastic layer, which, while 
forming the inner boundary of the cavernous tissue represents 
at the same time the inner insertion of the bundles of non-striped 
muscular cells mentioned above. 
The next point we have to consider is the constitution of the 
walls of Jacobson’s organ at a point a little further behind the 
one just described, 7. e. when the cartilage of Jacobson has 
become closed so as to form a complete capsule, such as is 
represented in fig. 4. The distribution of the different tissues 
as just described is very much the same, except that the epithe- 
lium of the lateral wall is separated from the elastic layer by a 
measurable stratum densely infiltrated with lymph-corpuscles. 
The elastic layer is very conspicuous. 
The cavernous layer, the distribution of the glands at the 
upper and lower sulcus, the sensory epithelium and the occur- 
rence of lymph-follicles, the presence of olfactory nerve trunks 
and plexuses of fine bundles of these in the median wall, is the 
same as in the parts anterior to this place. 
We have mentioned on a former page that past the region 
of the closed Jacobson’s cartilage capsule we again find this 
cartilage open in the upper part, assuming the shape of a 
trough, so that its cavity is in a free communication with the 
tissue at the side of the septal cartilage (see fig, 5), and we have 
also stated above that, beginning with this region, and through 
the greatest part of the crgan of Jacobson, the relations remain 
of this nature, with this difference, that as we pass farther back- 
wards the median or inner labium of the trough gradually increases 
in length. 
Now, with the change of the Jacobson’s cartilage from ak 
closed capsule into a trough-shaped plate, there occurs a very 
interesting change in the wall of the organ of Jacobson; the 
median wall, it is true, remains the same, and so does the dis- 
position of the glands of the upper and lower sulcus as well as that 
of the cavernous tissue and of the elastic layer in the lateral wall, 
but the subepithelial layer of the lateral wall alters so that this 
whole wall assumes a different aspect. Previously, we saw the 
epithelium of the lateral walls separated from the elastic layer by 
a thin layer of connective tissue densely infiltrated with lymph- 
corpuscles ; now, however, this subepithelial layer becomes greatly 
thickened, owing to the presence of serous glands, which oceupy 
the middle of the layer, so that this middle part of the lateral 
wall becomes changed into a fold forming a conspicuous projec- 
tion into the cavity of the organ of Jacobson. This fold may be 
therefore called the glandfold of the lateral wall. These glands 
