OEIGIN AND MOEPHOLOGY OF CHLOEOPHYLL COEPUSOLES. 239 



Dehnecke does not find to be correct ; he rather concludes that 

 the " etiolin bodies " are formed freely in the protoplasm, and 

 become green later. In his further conclusion, that assimilat- 

 ing and non- assimilating chlorophyll bodies are not different, 

 and that in many cases the same bodies may have successively 

 the function of storing starch and of assimilation, and vice 

 versa, his results coincide with those of Schimper. 



Up to this point, though the formation of the chlorophyll 

 bodies had been touched upon by various writers, still no clear 

 idea had been obtained of their ultimate origin; and though 

 it was well known and taught in all the text-books that chlo- 

 rophyll grains could multiply by division, still an origin by free 

 formation was ascribed to them, and it was compared with the 

 process of " free cell formation/' The way for a new view of 

 the ultimate origin of chlorophyll bodies and allied structures 

 was prepared by the results of the investigation of the nucleus 

 of the cell by various observers, but especially by Strasburger. 

 This author distinctly laid down the principle^ that there is no 

 free nuclear formation in embryo sacs, which were formerly 

 understood to supply the best examples of this process. He 

 further widened his generalisation so as to include all tissues, 

 and a translation of his own words will best convey his mean- 

 ing :" — " We are unable any longer to quote examples of free 



nuclear formation in the vegetable kingdom In all 



known cases in the vegetable kingdom cell-nuclei which make 

 their appearance are to be regarded as derivations of pre- 

 existing nuclei. The new nuclei result from a division of older 

 oues."^ 



This same character was found by Schimper^ to apply also to 

 bodies which we are now considering. His observations led 



1 'Bot. Zeit.,' 1879, p. 265. 



2 'ZeUbildung und Zelltheilung,' 3rd ed., 1880, p. 321. 



3 Observations recently made do not tally with this statement. Cf. K. 

 Prohaska, " Der Embryo-sac u. d. Endospermbiidung in der Gattung Daphne," 

 ♦ Bot. Zeit.,' 1883, No. 52. 



■* 'Bot. Centralbl.,' 1882, No. 44, which is a preliminary abstract of the 

 paper published, ' Bot. Zeit.,' 1883, Nos. 7—10. 



