74 EDWARD B. POULTON. 



fig. 1), and always situated at the base of the ganglion close 

 to the fibrous capsule. In vertical sections two to six cells 

 appear in one section. In a horizontal section across the base 

 of the ganglion about twelve cells are seen, and many others 

 also present appear to be belong to small ganglia in the 

 course of nerves entering the larger ganglion. The nerve- 

 cells are very large and fusiform, with branched ends (the 

 branches losing themselves among the nerve-fibres of the 

 ganglion). The large oval nucleus, containing a distinct 

 nucleolus, is coarsely granular in appearance, and does not 

 stain in hsematoxylin, although the finely granular cell-body 

 stains deeply. Large nerves with non-medullated fibres are 

 distinctly seen entering the base of the ganglion, and in a 

 deep horizontal section a ground-plan, as it were, of the gan- 

 glionic nerves is seen, as they radiate outwards from the base 

 of the ganglion. These nerves contain isolated nerve-cells, 

 and also small groups, in their course, the cells exactly resem- 

 bling those of the ganglion, except that they become polyhedral 

 when crowded together 



It thus appears almost certain that nerve-cells are inter- 

 calated in the course of sensory impulses from the periplieral 

 organs to the nervous centres. This is of interest in bringing 

 these terminations into closer connection with the related 

 terminal organs of sight and hearing, where ganglion cells 

 similarly intervene. If this be a true correlation, it seems 

 likely that ganglion cells will be found generally in the nervous 

 masses of the large gustatory papillee, now that attention is 

 directed to their existence. Microscopic ganglia on the nerve- 

 branches have been described, and a nerve is generally 

 figured in the axis of the papilla, but a true, large, compact 

 ganglion making up most of the papillary body is, I think, as 

 yet unmentioned. 



Beneath the epithelium containing the taste bulbs the 

 nerve-fibres (in the course of or between which there a])pear 

 to be many nuclei) are very numerous, and are cut in all direc- 

 tions owing to their irregular and sinuous course. There is 

 no doubt of their connection with the bulbs, but it would 



