4G8 EDWAIU) J!. I'OULTON. 



beneath the anterior areas. The taste-bulbs are, as far as I 

 have observed, entirely unique in being developed at the ends 

 of long papillary processes, up which the nerve-fibres can be 

 seen streaming from the central nervous mass, sometimes 

 accompanied by capillary blood-vessels. 



Sometimes the external surface shows indications of lobation, 

 the convexities corresponding to the bulbs ; but this is un- 

 common. In rare cases a papillary process may divide and end 

 in two bulbs. Gustatory pores are present, and are singularly 

 like those of normal bulbs. The outline of an exceptionally 

 long and distinct pore is given in fig. 12, from which its length 

 and diameter are easily calculated. The ordinary length of 

 the pores in this animal is not more than half that shown in 

 fig. 12. I never saw any protrusion of cells or processes from the 

 pore. There appear to be rather under 500 bulbs to the square 

 millimetre of surface. The bulbs are oval or fusiform, and their 

 sides rise gradually from those of the papiUary processes, of 

 which they are the expanded ends. Tiie structure of the wall 

 is of great importance in the organogeny of the gustatory termi- 

 nation. At the same time it was singularly difficult to be certain 

 as to interpretations, owing to changes that had taken place in 

 these delicate structures. However, after comparing immense 

 numbers of sections, I can confidently assert that many of the 

 elements are not modified epithelial cells, but are altogether 

 subepithelial in origin. There are seldom traces of the me- 

 ridional arrangement of cells that characterises ordinary bulbs. 

 The elements also differ in being packed loosely, and in being 

 very heterogeneous. 



In many bulbs I have detected the yellow stain that results 

 from the disintegration of blood in a capillary. Many of the 

 cells look as if they might have scaled off the sides of the oval 

 chamber, and thus have been added to elements intruded from 

 below. Such cells are fusiform in shape, sometimes thick, 

 sometimes attenuated, but they always stain difiierently from 

 the surrounding epithelium. Other cells have many processes 

 and resemble connective-tissue corpuscles, although they may 

 be nerve terminations. Others resemble small multipolar gan- 



