DEVELOPMENT OP THE CAPE SPECIES OP PERIPATUS. 497 



The external opening of the nephridium (fig. 37) is anterior to 

 the funnel. The course of the tube at this stage will be under- 

 stood by an inspection of figs. 37, 38, 38 a. It will be observed 

 from these that the nephridium — the part marked I. s. t. 3 in 

 fig. 38 — projects back as a tube which ends blindly (fig. 38 a, 

 I. s. t. S = sal. gl.). This backwardly projecting part gains an 

 enormous extension in the later stages, and is known in the 

 adult as the salivary gland. 



Kennel, who has correctly recognised the nephridial nature 

 of the salivary gland, has made what I cannot but regard as 

 one or two blunders in his description of its devevelopment. 



He says (No. 14, p. 38), " Von alien Theilen der friiheren 

 Segmentblase behalt nur der 3. Abschnitt, die blindsackartige 

 Ausstiilpung, ihren Charakter, indem dort die Zellen als 

 Epithel angeordnet sich erhalten ; er wird zum Trichter des 

 Segmentalorgans." Again, on p. 45, " Man findet ihn (seg- 

 mental trichter of salivary gland) an seiner alten Stelle, 

 allenfalls ein wenig weiter nach hinten verschoben, als kurzen 

 Blindsac, welcher von seiner Ansatzstelle aus schrag nach 

 vorn dicht am eigentlichen Kanal der Speicheldriise hin 

 verlauft, nwi aber nicht mehr mit Raumen des Lateralsinus 

 communicirt , sondern blind geschlossen ist ^' (my italics). 



Combining these passages with a statement on p. 38 as to 

 the breaking up of the Segmentblase of the oral papilla, it seems 

 clear that Kennel imagines (1) that the third somite does not 

 divide into a median and lateral part (though it does so in 

 other somites). (2) That the somite itself breaks up entirely 

 into a system of spaces, of which the lateral sinus is part. 

 This is implied by the first of the above quotations, and the 

 italicised parts of the second. It is also definitely stated on 

 p. 176 of Th. 1 (No. 13) in the following words: '' Letzere 

 (i.e. Segraentalhohle) werden spater ganzlich in den Lateral- 

 sinus und die Hohlung der Fiisschen umgewandelt und geben 

 den Hohlorganen daselbst, besouders den Segmentalorganen 

 z. Th. der Ursprung.'' (3) That the funnel of the salivary 

 gland becomes closed and persists in life as a vesicle. (4) 

 (from other statements on p. 39) that the funnel of the 



