322 ALFRED GIBBS BOURNE. 
of goblet-cells, looking very like those in the epidermis and 
probably secreting mucus.! 
The epithelial cells of the cesophagus are also many of them 
glandular, and the layer remains unchanged until we reach the 
gizzard. There the cells become very short, small, cylindrical 
cells, very closely set together; and here there is a cuticle and 
no glands. This cuticle is continuous from one end of the 
gizzard region to the other, but varies in thickness, becoming 
very thick in the middle of each gizzard and thinning out at 
either end, being thinnest in the cylindrical regions between 
the gizzards. 
In one segment behind the gizzard there are no glands in the 
epithelium, and then comes the region of tubular glands. In 
this region the epithelium on the surface appears only slightly 
glandular. In a section passing through the aperture of a 
tubular gland (fig. 24) the ordinary epithelium is seen to be 
continued some way down the follicle, and then its character 
gradually changes, the cells become broader and evidently 
glandular, but they do not become goblet-shaped, and they are 
all alike; their nuclei are smaller than those of the ordinary 
epithelial cells. ‘The follicles run straight down to the base- 
ment membrane, and then branch and coil. 
In the saccular region of the intestine the mucous membrane 
becomes thin again, and consists of a single layer of epithelial 
cells only. Most of these have granular contents and appear 
to be glandular, but they do not become much swollen, or at 
any rate are not so in any of my sections, which come from 
several different worms. 
In the rectal region the cells are all narrow and non- 
glandular. Longitudinal sections passing through the anus 
show that there is no sharp demarcation between rectal epi- 
thelium and epidermis. 
Of the special muscles of the pharynx and gizzards I have 
already spoken. In the wall of the rest of the cesophagus and 
of the intestine the usual muscles are present, a very thin layer 
1 These glands show best when the tissue has been prepared in Flemming’s 
mixture of chromic, osmic, and acetic acids. 
