BRANOHIAL SENSE ORGANS IN IOCHTHYOPSIDA. 135 
previously stated his conviction that the “ Geschmackorgane”’ 
or taste buds were the last remains of the skin sense bulbs of 
fishes, and Blaue now homologises the smell buds and the 
sense bulbs of the skin of fishes. 
But though he is convinced of this homology, he nowhere 
hints that the nose is to be regarded as a specialised portion 
of the so-called organs of the lateral line, and in fact accepts 
and supports Marshall’s gill theory of the nature of the nose, 
and derives his smell buds from skin sense bulbs which, 
originally present on the nasal visceral arch as in other cases, 
have wandered into the nasal cleft. 
Now, although sense bulbs are present on and along the vis- 
ceral arches of many fishes, they are not primitively there, 
their primitive position being above the cleft, not along it. 
Their presence along the arch is a later development. This 
fact and the facts of development as given before are entirely 
opposed to Blaue’s supposition. 
It is a curious commentary on the influence of the same set 
of facts on the views of different zoologists that while Blaue, as 
the result of his researches, advocates the gill nature of the 
nose, Prof. Wiedersheim, as he has kindly informed the writer 
by letter, since reading Blaue’s paper, considers it necessary, 
as most morphologists would, to give up entirely the notion 
that the nose is a gill-cleft. 
My own opinion does not rest on the researches of Blaue 
alone. Apart from those discoveries, I should believe myself 
justified in holding, as against the views of Prof. Dohrn and of 
my own teacher, Prof. Marshall, that the nose is the modified 
sense organ of a gill-cleft rather than a gill-cleft itself. 
But though maintaining that Blaue’s results are not neces- 
sary to support this view, yet, blending together those results 
and the facts recorded in this paper as to the development, 
&e., of the supra-branchial sense organs and of the nose itself, 
I believe that my view of the nature of the nose has so solid a 
foundation in facts that even the most sceptical zoologist can 
have little hesitation in accepting it. 
Shortly stated, the olfactory organ is a branchial sense 
