SPENGELIA, A NEW GENUS OF ENTEROPNEUSTA. 629 



" Piiuktsubstanz/^ accessory genital pores not perforating the 

 lougitudiual musculature, &c.) might almost be defined as a 

 Glaudiccps with synapticula. Spengcl specially mentions 

 the absence of synapticula as indicating the primitive character 

 of Glandiceps. In a recent paper in the ' Quarterly Journal 

 of Microscopical Science ' ^ dealing with impressions of 

 Ptychodera flava derived from an examination of fresh 

 material, I expressed the opinion that Ptychodera presented a 

 more primitive type of organisation than the other known 

 Enteropueusta. The discovery of this new genus, Spengelia, 

 which has so many points in common with Glandiceps (see 

 above), and yet which has synapticula between the branchial 

 bars and vestigial roots arising from the collar nerve-cord, goes 

 a long way to prove that Ptychodera is relatively primitive, 

 and that Glandiceps and Balanoglossus are derived forms. 



I hope to supplement the above account by a second part 

 containing an illustrated description of the anatomy of Spen- 

 gelia. What has been said is enough to establish the genus. 



My work, so far as it has gone on the preserved specimen, 

 has been carried out in Professor W. A. Haswell's laboratory 

 at the University of Sydney. I made some sections of 

 Ptychodera flava for purposes of control, but in addition I 

 have had the advantage of examining Mr. J. P. Hill's beautiful 

 preparations of three species of Ptychodera, besides having 

 his opinion on various points in my own preparations of Spen- 

 gelia, notably on the vestigial roots and the splanchnic layer 

 of '^Punktsubstanz." 



Sydney; May 10th, 1897. 



1 Vol. 40, p. 105. 



VOL. 40, PART 4. NEW SEE. X X 



