20 MARCUS M. HARTOG. 
VAUCHERIA is continuous and oogamous. The spermatan- 
gium is the distal end cut off from a short lateral tube. From 
the number of spermatozoa formed here, as compared with 
the vegetative nuclei of the ordinary protoplasm, it is certain 
that nuclear fission must precede spermatogeny; according to 
Berthold! there is here a formation of epiplasm comparable 
with that of the sporangia and gametangia of Cladophora. 
The “ oogonium ” is formed primitively as a lateral out- 
growth ; it contains at first numerous nuclei, the number of 
which is finally by fusion reduced to one.” A beak for the 
passage of the spermatozoa is formed as in Oedogonium, 
and the formative plasma undergoes mucous degeneration. 
Schmitz calls this mucified plasma of the beak a Richtungs- 
korper (polar body), and says it contains ‘‘numerous small 
nuclear fragments abstricted from the numerous nuclei of the 
young oogonium.”* If this meagre statement be accurate 
(and it is all we have), it would seem that nuclear divisions to 
form gametonuclei take place, and half the offspring pass into 
the epiplasm, and the other half fuse to form the pronucleus 
of the single oosphere—a process comparable with that of 
Peronospora (infra, p. 22). 
2. Colourless or Fungal Types. 
(a) Phycomycetes Zoosporez. 
It is convenient to consider separately the Phycomycetes 
with flagellate spores, comprising Chytridiex, Ancy- 
listee, Monoblepharis, Peronosporeex, and Sapro- 
legniex, as Zoosporee, in opposition to the higher group 
Aplanosporee, which never form organs of locomotion, and 
which comprises only Entomophthoree and Mucorini. 
ANCYLISTEH* are in appearance oogamous, the entire con- 
1 Op. cit., p. 305. 
2 According to Schmitz, ‘‘ Die Zellkerne der Thallophyten,” in ‘ Sitzungsb. 
d. Niederrh. Ges. zu Bonn,’ 1879. 
3 This is contained as a mere by-statement in a foot-note (on p. 225) to 
his paper, ‘“ Untersuchungen iiber die Befruchtung der Florideen,” in 
‘Sitzungsb. d. Berl. Akad.,’ 1883. A full account is much to be desired. 
4 My account of Ancylistew and Olpidiopsis is taken from Zopf, 
