300 W. BLAXLAND BENHAM. 
InteRNAL Anatomy, Pl. XVI, fig. 19. 
There are no especially strong septa (though those of 
xu/xiv, xtv/xv, and xv/xvi are slightly thicker than the 
rest), nor any great displacement of organs. 
The Alimentary Canal.—The pharynx, which occupies 
the Somites 111, rv, and part of v, is partially concealed by two 
pairs of lobulated masses, or “ salivary glands” (Sal.), lying 
dorsally and laterally in Somites 1v and v, the gland in the 
latter being bilobed. These are not modified nephridia, but 
consist of groups of large granular cells, deeply stained in 
borax carmine, and closely resembling the “salivary glands” 
of Eminodrilus,! &e. 
The pharynx is internally divisible into a ventral cuticulated 
region, and a dorsally placed flattened diverticulum, laterally 
produced, and lined by tall ciliated cells, as in many other, 
perhaps all earthworms. 
The cesophagus, which is not ciliated, is narrow and tubular: 
it passes backwards to the large sacculated intestine, which 
commences in Somite xv. 
The hinder region of the cesophagus in Somites x to x11I is 
highly vascular, and its wall thrown into folds; these become 
more definite in Somite xiv and give rise to a well-marked 
pair of calciferous glands. ach is a large sac, laterally 
placed, but extending dorsally and ventrally, so that the pair 
almost surround the cesophagus (Pl. XVI, fig. 20, cal.). Its 
structure calls for no remark. 
There is no gizzard to be detected on dissection, but in 
Somite vi the wall of the gut is slightly more muscular than 
elsewhere, the coat being about twice the thickness of that of 
the neighbouring tract, instead of eight to ten times as thick, 
as in the case of the functional gizzard in most earthworms. 
The sacculated intestine commences in Somite xvr, where it 
suddenly dilates, to become about three times the width of the 
cesophagus. This character it retains in Somites xVII, XVITI, 
and xix, beyond which it is spirally coiled (Pl. XVI, 
figs. 19, 21). This condition has been described in Didy- 
' Benham, ‘Journ. R. Micr. Soc.,’ 1891, pl. iv, fig. 12. 
