202 W. WALDEYER. 
SS 
figure, as long rods (Tonnenform, couronne a batonets of 
Carnoy), then follows the transverse division of these rods at 
the equator, and the separation of the two halves towards the 
daughter-poles. This is quite at variance with all previous 
conclusions. According to this statement the chromatic figure 
behaves just like the spindle figure. 
According to Carnoy, too, a karyokinetic arrangement of 
chromatic threads at the two poles can occur without any lon- 
gitudinal or equatorial transverse splitting, e.g. loc. cit., pl. v, 
fig. 178, in Clubiona (an Arachnid). A. Bolles Lee (32) has 
lately made the same statement for karyokinesis in the sper- 
matogenesis of the Chetognatha. No longitudinal splitting 
of the equatorial chromatic elements (chromosomes) was seen 
in Forficula by La Valette St. George; on the contrary, his 
figures show a transverse division (see Festschrift fiir A. v. 
Kolliker, Taf. i, figs. 35, 36, &c.). Prenant lhkewise makes 
similar statements. In all such processes equal distribution of 
the chromatic substance would be uncertain. 
As we have already mentioned, Carnoy found that the 
longitudinal splitting, which he in nowise denies, may even 
occur during the stage of the formation of the daughter-nuclei. 
If this is true (Ascaris megalocephala, acccording to 
Carnoy), longitudinal splitting is of only a very subordinate 
value. Flemming (68) has recently taken up a determined 
position against this reduction of longitudinal splitting to a phe- 
nomenon of subordinate significance. He found in the sperma- 
togenesis of Salamander an abnormal form, which appears 
at first sight to confirm Carnoy’s assertion. But here longi- 
tudinal splitting of the threads occurs, at the commencement 
of karyokinesis, in the so-called spirem stage (see my previous 
figure). The two sister-threads remain connected so as to form 
an elongated ellipsoid, stretch themselves, and all the ellipsoids, 
greatly elongated, now arrange themselves ina “ barrel figure” 
(couronne a batonets of Carnoy) around the equator of the 
spindle figure. The chromatic elements (Faden, chromo- 
somes) are, however, not simple primary rods, but ellipsoid 
rings, greatly drawn out, each of which consists of two sister- 
