KARYOKINESIS AND ITS RELATION TO FERTILIZATION. 213 
Carnoy uses the terms : 
Spirem = 1. Scission du boyau ou du peloton. 
Aster = 2. Couronne équatoriale. 
Metakinesis = 8. Ascension polaire. 
Dyaster = 4. Couronnes polaires. 
Dispirem = 5. Reconstitution du noyau. 
It is, however, desirable that in the classification of the 
mitotic processes in different stages the most important of 
them—the longitudinal splitting of the segments, and 
the appearance of the nuclear spindle,—should have separate 
places. They may perhaps be grouped as follows, so as to 
obtain a classification suitable to all cases: 
A, I. Resting mother-nucleus. 
II. Mother-coil (spirem). 
III. Splitting of the loops and spindle-stage. 
IV. Mother-star (monaster). 
B. V. Metakinesis. 
C. VI. Daughter-stars (dyaster). 
VII. Daughter-coils (dispirem). 
VIII. Resting daughter-nuclei. 
A couple of words must be added in explanation of this 
suggested grouping. 
All workers agree that the resting stage is followed by 
another, which shows the chromatin arranged in the form of 
distinct threads. Like the words “ Knauelstadium ” or “ Mut- 
terknauel” (skein-stage, mother-skein), “‘ Spirem” does not 
prejudge anything as to whether there is one thread only 
or numerous threads, which at first appear inextricably 
intertwined. It does not appear to me to be necessary to 
speak of a denser or a looser coil or skein. On the other 
hand, it is incontestably very important that in the further 
course of mitosis distinctly separate segments (chromo- 
somes) in the form of rods or loops arise; that these 
undergo a longitudinal splitting, and that the nu- 
clear spindle figure makes its appearance at this stage. 
