CONSTITUTION AND DEVELOPMENT OF TERMITES. 301 
A? of Stage 1v; c® and p® from the forms with wing-buds of 
Stage tv; and lastly, u* always from form c? of Stage rv. 
To this fifth group belong further certain individuals 
which I regarded as abnormalities before I ‘was acquainted 
with the facts I have recorded about the soldiers of Calo- 
termes. 
They are nymph-soldiers, or nymphs with the buccal ap- 
paratus of soldiers. They probably lose their wings and 
become simple soldiers, as do those of Calotermes.! 
VI. Perfect insects—that is, with fully developed 
wings and capable of flight (Pl. 17, fig. 11). The number 
of antennal joints remains, as in the nymphs, at seventeen or 
eighteen, but they are always entirely pilose. These specimens 
are distinguished by a general piceous colour, save for the 
mouth parts, tarsi, and apices of the tibie, which are testa- 
ceous. They originate from the nymphs of the first form. 
VII. Complementary (Pl. 17, figs. 16—18, 21) or sub- 
stitute (ad., figs. 15,22, 25) royal forms, sexually ma- 
ture, or nearly so, but in the guise of the larva or 
nymph, or else resembling an imago, partly infuscate 
and with the wings torn. 
The latter spring from immature, not fully darkened perfect 
insects ; the others, in the larva or nymph form, arise from 
the royal larve 2n® of group V, or from nymphs of the second 
form, or finally from nymphs of the first form (fig. 23). 
The royal forms derived from nymphs of the first form or 
from not fully darkened perfect insects are all substitute, 
never complemental forms. 
The wing-rudiments may be wanting, or more or less short. 
The body may be of the whitish yellow of old paper, or may be 
more or less extensively blotched with brown. The abdomen 
is much inflated, so that these forms are generally recognisable 
at a glance. 
They exhibit a further characteristic in the possession of 
‘ It now seems to me probable that they should be compared with the 
egg-hatching workers of the honey-bee.—G. B. Grassi, October, 1896. 
