﻿ON 
  A 
  VKW 
  SPEOIES 
  OF 
  THE 
  GENUS 
  HAPLOTAXIS. 
  315 
  

  

  Bergli 
  (8), 
  in 
  1886, 
  describes 
  in 
  detail 
  the 
  development 
  of 
  

   the 
  funnels 
  in 
  Lumbricns, 
  but 
  he 
  failed 
  to 
  trace 
  the 
  deve- 
  

   lopment 
  of 
  the 
  genital 
  ducts 
  therefrom. 
  The 
  genital 
  funnel 
  

   develops 
  as 
  a 
  thickening 
  of 
  the 
  peritoneal 
  cells 
  immediately 
  

   above 
  (dorsad 
  of) 
  the 
  nephridial 
  funnel 
  (which 
  of 
  course 
  

   does 
  not 
  disappear 
  in 
  this 
  earthworm) 
  ; 
  the 
  development 
  in- 
  

   volves 
  partly 
  the 
  cells 
  forming 
  a 
  covering 
  for 
  the 
  back 
  of 
  the 
  

   nephridial 
  funnel 
  and 
  paitly 
  the 
  cells 
  forming 
  the 
  anterior 
  face 
  

   of 
  the 
  septum 
  itself. 
  The 
  same 
  is 
  true 
  both 
  for 
  oviduct 
  and 
  

   sperm-duct, 
  and 
  this 
  very 
  close 
  association 
  of 
  nephridial 
  

   fnnnel 
  and 
  coelomostome 
  is, 
  it 
  seems 
  to 
  me, 
  likely 
  to 
  be 
  of 
  

   interest 
  when 
  the 
  development 
  of 
  the 
  entire 
  nephridial 
  funnel 
  

   is 
  fully 
  known. 
  

  

  At 
  present 
  we 
  do 
  not 
  know 
  whether 
  the 
  " 
  marginal 
  " 
  cells 
  

   of 
  the 
  Lumbricid 
  nephridial 
  funnel 
  are 
  coelomic 
  in 
  origin. 
  

   It 
  is 
  quite 
  possible 
  that 
  they 
  are. 
  For 
  the 
  '^central 
  celP' 
  is 
  

   probably 
  the 
  original 
  funnel-cell, 
  which, 
  according 
  to 
  Vej- 
  

   dovsky, 
  divides 
  so 
  as 
  to 
  permit 
  the 
  tubule 
  to 
  communicate 
  

   with 
  the 
  coelom. 
  If 
  it 
  should 
  turn 
  out 
  that 
  these 
  peripheral 
  

   marginal-cells 
  are 
  coelomic, 
  then 
  the 
  close 
  topographical 
  rela- 
  

   tion 
  of 
  genital 
  funnel 
  with 
  the 
  nephridial 
  funnel 
  described 
  and 
  

   figured 
  by 
  Bergh 
  will 
  indicate 
  that 
  the 
  whole 
  '' 
  nephridium 
  " 
  

   of 
  earthworms 
  is 
  a 
  "nephromixium." 
  

  

  A 
  wholly 
  different 
  history 
  is 
  given 
  by 
  Roule 
  (13), 
  in 
  1889, 
  

   for 
  an 
  Enchytraeid. 
  In 
  the 
  earlier 
  stages 
  in 
  the 
  development 
  

   of 
  " 
  Enchytrseoides 
  marioni" 
  the 
  twelfth 
  segment, 
  in 
  

   which 
  at 
  a 
  later 
  period 
  the 
  sperm-ducts 
  will 
  arise, 
  contains 
  

   no 
  excretory 
  organs, 
  though 
  these 
  are 
  present 
  in 
  the 
  pre- 
  

   ceding 
  segments, 
  IX, 
  X, 
  Xf, 
  and 
  in 
  the 
  segments 
  following 
  

   it, 
  namely, 
  XIII, 
  XIV, 
  etc. 
  The 
  young 
  sperm-duct, 
  when 
  it 
  

   does 
  ultimately 
  make 
  its 
  appearance, 
  is 
  ''en 
  tout 
  semblable 
  

   a 
  une 
  tres 
  jeune 
  nephridie, 
  et 
  deplus, 
  il 
  occupe 
  exactement 
  

   la 
  place 
  qu'aurait 
  Torgane 
  segmentaire 
  s'il 
  s'etait 
  developpe 
  

   dans 
  la 
  Xllme 
  anneau." 
  Its 
  mode 
  of 
  origin 
  is 
  stated 
  to 
  be 
  

   quite 
  like 
  that 
  of 
  a 
  nephridium, 
  after 
  this 
  has 
  separated 
  

   itself 
  from 
  the 
  nephric 
  cord 
  (which 
  is 
  observable 
  in 
  the 
  pos- 
  

   terior, 
  but 
  not 
  in 
  the 
  anterior 
  segments) 
  ; 
  it 
  now 
  consists 
  of 
  

  

  