516 EIOHARD EVANS. 



glands open by tlie sides of the nerve cords, near the 

 posterior end, that is, they open almost exactly where we 

 should expect a renal organ to open, and so far this is in 

 favour of their homology with the renal organs ; but we 

 know nothing of their development. It is not known 

 whether they belong to the anal segment, as in P. Ed- 

 wardsii, or to another segment situated between the anal 

 and the genital segment, which in P. No vaj-Zealandias 

 has lost all traces of its appendages. If they belong to the 

 former there is a very close relation between them and the 

 anal glands of P. Edwardsii, but if they belong to the 

 latter, the most that can be said is that they are thus homolo- 

 gous in the same sense that the renal organs of the various 

 segments are homologous with them. 



In P. Leuckarti the ^'accessory glands " differ from those 

 of P. N ova3-Zealand ia3 only as regards their external open- 

 ings, which are situated close together near the middle line, 

 between the genital orifice and the anus — a position reached 

 by a very slight amount of shifting towards the mid-venti-al 

 line. It seems probable, when other genera are taken into 

 consideration, that in the above two species which belong to 

 the genus Peripatoides, the "accessory glands" are de- 

 rived from the anal somite. 



The condition existing in P. Leuckarti leads naturally to 

 that found in Eoperipatus, where the accessory glands open 

 into a common pit situated in the mid- ventral line between the 

 genital pore and the anus; that is, the external opening is 

 placed exactly where we should expect to find it if the acces- 

 sory glands belonged to the somite of the last leg-bearing 

 segment. But if the accessory glands of Eoperipatus are 

 homologous with the renal organs, they cannot belong to the 

 last leg-bearing segment; for in both male and female there 

 exists a well-developed renal organ in that segment, though 

 in the adult male the external portion of the duct is not 

 found. It follows that the accessory glands of Eoperi- 

 patus must be either the renal organs of the anal segment 

 as in P. Edwardsii, or the crural glands of the last leg- 



