TWO NKW SPECIES OF ONYOHOPidORA . 5f9 



passuge it seems absolutely clear that the somites behind the 

 genital segment do not give rise to the male accessory glands 

 of P. capensis. It follows from this that the glands in 

 question cannot in any way be described as modified renal 

 oi-gans, unless the genital somite is capable of giving rise to 

 two sets of similar organs, one of which is modified as the 

 genital ducts, and the othei- as the male accessory glands — 

 an absolutely gratuitous supposition, for tlie body segments 

 of the Peripatidas, as far as is known, never give rise to 

 more than one renal organ or its modified homologue. It 

 seems, therefore, that the " obvious homology " of the male 

 accessory glands of P. capensis with the anal glands of P. 

 Edwardsii, the pygidial glands of P. No va3-Bri taunia?, 

 and the accessory glands of P. No vae-ZealandiiV, P. 

 Leuckarti, and Eoperipatus, is far from being proved, and 

 that the possibility of their being merely modified crural 

 glands must not be lost sight of. It may be that, when we 

 shall have learnt more about the origin of these glands in 

 the different genera, they will be shown to be homologous ; 

 but for the present we must at least suspend judgment, as 

 the formation of a premature conclusion tends to obscure the 

 problem which has to be solved, and the view that they 

 are homologous is still sub judice. 



(/() The Crural Glands. — The occurrence of these glands 

 lias already been mentioned in describing the position of their 

 external apertures. There are four pairs of them ; two in 

 each of the first and second prjcgeuital pairs of legs. They 

 are tubular glands which almost exactly equal in size 

 the renal organs of the segments in which they occur 

 (figs. 27 and 28). They are lined internally with a thin 

 chitinous layei', under which are found the shoi't columnar 

 cells, which are supported by a thin muscular coat and a 

 peritoneal investment. Tlie occurrence of two crr.ral glands, 

 in each of the legs in which they occur, is a feature in which 

 Eoperipatus Horsti resembles P. Edwardsii. Itappears, 

 however, that they occur in a far greater number of legs in 

 the latter than iu ihe former (7). 



