574 W. BLAXLAND BENHAM. 



seems to be a droplet of some fluid, but not of an oily 

 character; there is no marked refriugency, and reagents 

 point to different substance. 



The " granules/' which are somewhat greenish in colour, 

 are contained within the vacuoles, as can most certainly be 

 recognised in crushed cells (see fig. 6, a), and each granule 

 exhibits " Brownian movement" therein, whether the vacuole 

 be still within its cell or isolated. As a rule each vacuole 

 contains a small granule, and never more than one, but 

 frequently the vacuole contains none. I have not seen any 

 granule independent of a vacuole; the two are genetically 

 related, but whether the granule is formed within the 

 vacuole, or the fluid of the vacuole arises as a result of solu- 

 tion of the granule I cannot determine. 



These vacuoles are fairly constant in size, but the granules 

 vary Avithin small limits and in diffei'ent cells, while the 

 number in different cells is also subject to considerable 

 variations. The resemblance in size that a vacuole bears to 

 a globule naturally suggests some relation between the two, 

 but, as will be seen below, there is a chemical difference 

 between these things, though it seems probable that there is 

 a genetic bond connecting them in a series. 



I have already mentioned that most of the eleocytes 

 contain, also, a few " granules " in vacuoles ; sometimes the 

 characteristics of the two cells, which for convenience I refer 

 to by separate names, are united in a single cell (see fig. 7). 

 There is therefore little doubt as to the relation of one to the 

 other, and in some specimens of Octochaetus the resem- 

 blance is still closer, in that some of the " lamprocytes " 

 contain " vacuoles " which are without granules. 



We thus have, if we regard the eleocytes and lamprocytes 

 as derivatives one from the other, four conditions : 



(a) Cells containing nothing but oily globules. 



{h) Cells chiefly with globules, with few or many granules 

 in vacuoles. 



(c) Cells with only one granule in each vacuole. 



(d) Cells with vacuoles only. 



