A NEW SPECIES OP MOiNILIGASTER FROM INDIA. 371 



middle of the cell (fig. 10) ; outside this is a thick coating 

 of muscles in various directions, with of course numerous 

 blood-vessels. But after a short course this simple epithelium 

 gives way to a glandular coat (fig. 11), recalling the structure 

 of the " prostate." 



The '^ prostate" or glandular wall of the atrium (fig. 9), 

 however, is lined by a layer of tall, thin, glandular cells, with 

 nuclei (? at any rate deeply stained structures) placed at the 

 sides of the cells, as if compressed ; somewhat similar to, 

 though much narrower than, the epithelium figured and 

 described by Beddard. 



The muscular coat, consisting of circular, longitudinal, and 

 oblique fibres, does not immediately surround this epi- 

 thelium as in M. Barwelli, but the two are separated by a 

 considerable area, about twice as deep as the epithelium itself, 

 which appears to be occupied partly by certain longer cells 

 {gl.) than the lining epithelium, and partly by the necks of 

 the extra-muscular multicellular glands (see figs. 8, 9). 



In a foot-note on p. 573 of his memoir on Ocnerodrilus/ 

 Beddard refers to the fact, as observed in M. Barwelli, that 

 there is " hardly any distinction between a glandular and 

 non-glandular section of the atrium." In the present species, 

 as is shown, there is such a distinction very well marked. 



Outside the muscular coat are the groups of glandular cells 

 figured by Beddard. Each group (fig. 9) consists in a section 

 of some eight to ten club-shaped cells, the narrow necks of 

 which pass through the muscular coat, and can be 

 traced in bundles nearly up to the epithelium. Each 

 group is, in fact, a multicellular gland. This arrange- 

 ment is quite evident in my sections, and the necks or ducts 

 of the glands are quite noticeable, though I have been unable 

 to trace the necks right up to the epithelium ; the latter, how- 

 ever, is traversed by narrow clear tubes, resembling ducts 

 and which differ from the epithelial cells in having no inner 

 boundary (as at a, fig. 9). The nucleus of each cell of the 

 multicellular gland is naturally at the bottom, the widest part 

 ' • Trans. Roy. Soc. Edin.,' xxxvi, part 2. 



