584 FRANK E. BEDDAED. 



integumental network, which must, therefore, be regarded as 

 secondary, but the anterior nephridia at any rate are con- 

 nected on each side by a continuous longitudinal duct lying 

 within the coelom. 



(2) In the young worm the reproductive organs agree with 

 these organs in other earthworms; in the" adult, a large 

 unpaired sac lying over the gut is developed ; this sac encloses 

 the receptacula ovorum, and opens by a median pore on Seg- 

 ment 13. It is developed from mesoblastic tissues, and is not 

 therefore the morphological equivalent of the spermathecse in 

 Lumbricus, &c., but it performs the same function j the sac 

 is formed internally and then grows out towards the epi- 

 dermis ; it is at first in open communication with the coelom ; 

 its front wall is formed out of the intersegmental septum 

 between Segments 13, 13 ; the ovaries are enclosed by it, but 

 disappear early, before the sac is completed ; otherwise the 

 ova would be probably unable to enter the egg- sac which 

 becomes nearly completely shut off from the sac ; the two are 

 in communication only by the oviducal funnel, which has 

 become divided by the growth of the spermathecal sac into 

 two separate tubes, one opening into the spermathecal sac, the 

 other into the closed egg-sac ; they unite, of course, to form 

 the oviduct itself, which opens on to the 15th segment, 

 reckoning by the external furrow, but on to the border line 

 between Segments 14, 15, reckoning by the septa. 



(3) The testes and the vas deferens funnels are quite typical 

 in their structure and position ; so, too, are the (two) pairs of 

 sperm-sacs (in Segments 11, 12). The sperm ducts are not, 

 as they are in other Eudrilidse, dilated to form sperm reser- 

 voirs ; they open into tubular atria, with thick muscular walls 

 and glandular lining, near to their blind extremities; the two 

 atria open by a common pore upon the border line between 

 Segments 17, 18 ; each is furnished with a short penial seta 

 not ornamented. 



(4) The alimentary tract has no calciferous glands or 

 ventral oesophageal pouches such as are found in other Eudri- 

 lidee; at the end of the oesophagus are three gizzards, one to a 



