288 J. #, S; MOORE. 
fine achromatic line stretched round the nuclear membrane. 
My preparations indicate both these methods of procedure. 
The protoplasmic contents of the cell become radially dis- 
posed, not directly to the centrosomes, as in the divisions of 
the previous spermatogenetic period, but towards the outer 
surface of the daughter-archoplasms (fig. 43, 7.), and it conse- 
quently follows that the sphere of the first period is structurally 
less complex than that of the second. In cells which possess 
an archoplasm, any radiation in the cytoplasm external to this 
structure has not generally been considered a part of the 
attraction sphere; neverthelesss, such external radiations are 
obviously similar to those directly related to the centrosomes 
in the cells of the first spermatogenetic period, where they 
would certainly be regarded as a portion of the sphere. To 
save confusion, therefore, I shall speak here, as I have done 
elsewhere, of spheres which possess an archoplasm, as compound, 
and those which do not, as simple, and thus avoid the neces- 
sity of determining whether any particular set of radiations 
should or should not be regarded as constituents of the 
sphere. 
26. The ring chromosomes, which, when fully formed, 
become dispersed over the nuclear periphery, like those of the 
first spermatogenetic period, are in like manner connected to 
one another by numerous filamentous strands of linin (fig. 
43, 1.). 
The nuclear membrane eventually becomes irregular, and, 
giving way at various points, leaves the chromosomes to collect, 
by the contraction of these connecting filaments, into a long 
oval mass stretched across the nuclear sap between the centro- 
somes (fig. 44). The nuclear sap is traversed from the first by 
numerous fine strands, putting the chromosomes into connection 
with the outer cytoplasmic network, and which are in all pro- 
bability part of the latter, dragged inwards from without after 
the disruption of the nuclear wall. The central chromatic 
mass 1s somewhat stretched, and more firmly attached to 
the old nuclear surface in the direction of the spheres, appear- 
ing as if slung between the centrosomes (fig. 44). 
