294 Jo Bea S. MOORE: 
(figs. 75 and 76), the same formation of a nuclear vacuole in 
each daughter-cell (fig. 76, v.v.), the same gradually diminish- 
ing connecting spindle filament between the nuclei (figs. 76 
and 77), and lastly, the same formation of nuclear grooves 
(fig. 77, m.g.) along which the spheres travel to the equatorial 
side. Further, when the spheres have reached some point 
halfway between the polar and equatorial nuclear faces, the 
archoplasm leaves the nuclear wall. The centrosomes (fig. 
79, c.) pass on to the outer archoplasmic surface, and from this 
there passes a fine protoplasmic strand (fig. 80, f.) to the cell 
periphery, and the cell membrane is indented where this 
_ strand perforates it as the whiplash-like spermatozoon tail. It 
thus becomes evident that the metamorphoses described in 
§ § 22, 35 are nothing more nor less than abortive attempts 
at tail-formation, and it consequently follows that the 
synaptic phase in these fishes marks the assumption 
by the cells, during spermatogenesis, of a flagellate 
condition. 
40. Besides the nucleus, there appears a dusky condensation 
of the cytoplasm (#., figs. 80—83), which at first sight gives the 
cells the appearance of possessing more than one attraction 
sphere, and is obviously similar to the nebenkern of the 
preceding generation (of § 36). This body when it first appears 
is closer to the nucleus than the archoplasm (fig. 80), and 
in the latter there is seen at the point of origin of the 
flagellum a clear round vesicle (fig. 82, a. v.), which enlarges 
and eventually moves, with its archoplasmic surroundings and 
the centrosomes, into close apposition with the nucleus 
(fig. 83, a. v.). 
41, The nuclear chromatin rises up into a shallow collar 
round the base of the archoplasmic vesicle, while the rest 
of the chromatic substance, contracting from the nuclear 
membrane, becomes condensed into a flask-shaped mass below 
the collar (figs. 84 and 85). This contraction increases rapidly 
while the collar, elongating, spreads into the nuclear membrane 
at the base of the archoplasmic vesicle, to form a small chro- 
matic flange round the neck of a_ bottle-like structure 
