29, ARTHUR BOLLES LEE 
To ascertain this we must return to the study of the earlier 
telophase, or polar clump. In the daughter-star of the 
anaphase (figs. 8, 4, 5, 61) we have a loose assemblage of 
chromosomes, radially arranged in a ring. These contract into 
short staves; and as they contract the whole figure shrinks 
(figs. 29 to 34), so that the staves become closely huddled 
together and come into contact by them margins. ‘They 
generally seem to agglutinate there, and their outlines become 
hardly distinguishable, indeed very often quite indistinguish- 
able. The clump then appears (figs. 30, 31, 33) as an almost 
homogeneous ribbed disk, with a central pore, generally 
obturated by a perforated membrane or web formed (as shown 
by profile views) by the confluent remains of the polar spindle 
fibres. The mutual contact or agglutination of the chromosome 
staves takes place first in the region of the clump that is nearest 
to the pole, their more distal portions remaming longer free : 
so that at this stage we get the image of a compact ring with 
digitiform processes depending from it—the ‘ figures pectini- 
formes’ of Henneguy (figs. 32 and 34). In badly fixed cells 
the clumping results in a formless mass, in which the chromo- 
somes seem to have become completely fused together. This 
state is shown in fig. 834. But, as I gather from the study of my 
most favourably fixed specimens, this is an artefact ; and there 
is not at any time a real fusion of the chromosomes, but only 
intimate contact to the point of indistinctness, or possibly 
superficial agglutination.’ Fig. 83 seems to me to show the 
utmost degree of agglutination that should be taken to be 
normal; and the real state of things to be fairly well repre- 
sented by fig. 30 or 31. 
Careful examination of the staves of the clump at this 
stage seems to show that they are always in reality double 
structures ; for in favourable cases they show unmistakable 
indications of a longitudinal duplicity. In fig. 29 there are 
four staves, marked with a cross, which show this. In the left- 
hand one (near the top) the tip is distinetly bifid ; and this is 
1 Cf. Janssens, ‘La Cellule’, xix. 2, 1901, p. 86, and Janssens et 
Dumez, ibid., xx.2, 1908, p. 450 and fig. 15, who have arrived at the same 
conclusion. 
