STRUCTURE OF CERTAIN CHROMOSOMES 23 
also the case with the one at the bottom. In the two right-hand 
ones the tips are distinctly double ; and by careful focusing 
it can be made out that each of these staves is composed of two 
longitudinal moieties, superposed and to a slight extent twisted 
round one another. And in three or four of the short dark 
staves of the inner tier there can be seen a light longitudinal 
dividing line (not sufficiently clear in the drawing). 
In fig. 83 nearly one-half of the twenty-one staves drawn are 
seen to be notched at the periphery, and two of them show a 
longitudinal dividing line continuing the notch inwards. In 
fig. 30 I find three cases similar to these, and in fig. 82 two. 
Thave no doubt that with better fixation these nuclei would have 
shown several more such cases. In the clump of fig. 31 I think 
I can detect three or four similar cases, though doubtfully. 
The clump does not long remain in this state of dense ag- 
glomeration, but soon begins to expand into the telophasic 
ring. The manner of this expansion is as follows. Amongst 
the staves of the clump—but never on their outer surfaces— 
there appear certain hyaline globules which, growing, push the 
staves apart and so loosen the clump. In fig. 88 are shown two 
such globules, one to the right, and one to the left ; in fig. 35 
thiee (on the left ; one very indistinct) ; m fig. 87 five ; in the 
nucleus of fig. 86 there are a dozen or so, of which only a portion 
of one (at the left) could be shown in the drawing, the rest being 
too much masked by the sheaths. In fig. 62, to the right, are 
seen three ; in fig. 67 two can just be glimpsed (at the left and 
middle). These globules are entirely hyaline and uncolourable. 
Their outlines are generally quite smooth. They are, as I think, 
ovoid in shape, not spherical : they may show a circular outline, 
as in the left-hand ones of figs. 88 and 43, and other places ; but 
that is the expression of a transverse section of them. I suspect 
that there is formed at first one of them for each chromosome. 
If that be the case it is a likely hypothesis that they consist of 
the clear contents of the sheaths of the chromosomes, expressed 
from them by the pressure of the clump. But it is difficult to 
ascertain the number formed, because they soon fuse with one 
another into a small number of large globules, see figs. 48, 44, 46. 
