STRUCTURE OF CERTAIN CHROMOSOMES a 
pole. They consist, all of them, of tightly-folded V's, appearing 
as short staves with the spindle-fibre insertion at the end. But 
they are certainly folded V’s with the insertion at the apex: 
the two limbs can be made out with certainty at the tips of 
four of them; and a longitudinal duplicity can be at least 
climpsed in all of them.! I find the same state of things exactly 
in Oedipoda germanica, Oe. coerulescens, Oe. 
(Mecostethus) parapleura, Gomphocerus rufus, 
Stenobothrus morio, St. biguttulus, and some 
other species of § t eno b othrus which could not be determined 
with certainty. Sothat in all the Acrididae I have examined 
the folding takes place not later than the early anaphase. And 
as at this stage the images are not obscured by the crowding 
of the chromosomes which takes place in the polar clump, there 
can be no doubt about the folding actually occurring. 
So also in the Locustidae. Fig. 64 shows an anaphase of 
a spermatogonium of Decticus verrucivorus. The 
chromosomes are here smaller than in the Acrididae, and appear 
for the most part as short rods with the spindle-insertion at the 
end. But it can be made out in favourable instances that they 
are in reality folded V’s ; and where this cannot be done, the 
analogy with those of the Acrididae puts it out of doubt that 
they are in the same case. Similar images are afforded by 
Decticus griseus, Locusta viridissima, L. can- 
tans, and Pterolepis aptera. In Gryllotalpa 
vulgaris and Gryllus campestris I find apparently 
the same state of things, the anaphase chromosomes (with 
the exception of the monosome in Gryllus) appearing 
as short rods inserted by one end on the spimdle. These 
apparent rods are too small to be analysed with certainty ; but 
judging by the analogy of those of the other Orthoptera 
mentioned there can be no doubt that they are in reality 
1 The drawings figs. 12 and 13 (Dissosteira carolina), and 18 
(Steiroxys), of the paperof Davis, “Spermatogenesis in Acrididae’’, 
in ‘Bull. Mus, Comp. Zool. Harvard’, with the interpretations given, 
pp. 69, 70, 71 of the text, should, as I conceive, be corrected in the 
sense indicated above. 
