50 D. KEILIN 
methods were already advocated by Miiller (1895), Zimmermann 
(1898), and Retzius himself, for the detection of minute, or even 
intracellular, capillary ductules of secretion. 
(d) Septal glands. 
The salivary gland cells in all earthworms are intimately 
connected with some other cell aggregates which, being cyto- 
TEXT-FIG. 5. 
A and B. Sections of the ciliated pharyngeal epithelium of Lum- 
bricus sp. (fixed in Champy’s solution and stained with Iron- 
haematoxylin) demonstrating that the clubbed nerve fibrillae 
of Retzius are the intra-epithelial mucin ductules (d/. mu.) with 
their discharge pockets (d. p.); ¢. = cilia; mu. = contracted mucin 
in some of the discharge pockets. A x 734; B x 734. 
logically similar to the salivary cells, differ from the latter m the 
fact that they are completely devoid of mucin (PI. 3, fig. 2, e. gl.). 
Similar glandular aggregates, devoid of mucin, are found 
posteriorly in the coelomic cavity, surrounding the oesophagus. 
In places I believe that I have been able to trace a communica- 
tion between these deeply-lying glandular elements (septal 
glands) and the pharyngeal bulb. In other places, although I 
