THE PROTOZOA PARASITIC IN ARCHOTERMOPSIS WROUGHTONI 253 
Anterior Organ (Centroblepharoplast). 
Pseudotrichonympha pristina terminates at the 
anterior end in the curious organ found in the Tricho- 
nymphidae and described under various names by different 
observers: thus the Italian workers designate it as ‘la botti- 
chia’, “il cappuccio’, or ‘il mammillare’; to it Hartmann has 
applied the term ‘ Kopforgan’, and Porter * the nipple-like 
part’. Recently, however, Kofoid and Swezy have identified 
it as a centroblepharoplast, the name which I prefer to adopt. 
In P. pristina it is composed of two portions, an inner 
tube-like one surrounded by a sheath which appears to cover 
it completely (Pl. 10, figs. 3, 5). This ectoplasmic sheath at 
its distal extremity becomes continuous with the rest of the 
body, and this is the only region where differentiated ectoplasm 
is found. I have been unable to detect any trace of a break 
in the tip of the sheath such as one would expect were the 
inner region in reality a tube capable of expelling or taking in 
liquids as some observers would have us believe. Each anterior 
flagellum takes origin from a granule situated on the inner surface 
of the ectoplasmic layer of the centroblepharoplast. These 
granules are difficult to detect, but in a few suitable prepara- 
tions they are unmistakably demonstrated (Pl. 10, fig. 5, B.G.). 
Finally, from the extreme end of the organ there arise two 
fine threads, which, taking a parallel course down the centre 
of the endoplasm, diverge at thei distal ends to reach the 
nuclear membrane where they are attached (Pl. 10, figs. 38, 
5, s.7.). It seems indubitable that there is such attachment, 
for in specimens whose nuclei have been thrust out of position 
the threads are still seen running to the membrane. Thus 
the nucleus is more or less fixed in position by these threads, 
in contrast to the * free’ nucleus described by Grassi. 
Striations and Granules. 
The striations that are seen crossing the body in a longitu- 
dinal series arise from the centroblepharoplast. They consist 
of ridges in the body surface, and thus broadly agree with those 
