CYTOPLASMIC INCLUSIONS OF THE GERM-CELLS OTT 
more than one-half the size of the same structures in the 
spermatocyte, it is difficult to obtain satisfactory evidence of 
any increase in size of the individual mitochondria. 
With reference to the sorting out of the Golgi elements or 
dictyosomes during the maturation divisions, attention is 
drawn to recent work on Limax agrestis, where it has 
been demonstrated that the number of dictyosomes in the 
spermatocyte is eight, and in the spermatid two (18). In 
all probability, though no count is possible in Cavia, the 
number of platelets or dictyosomes in the spermatid is approxi- 
mately one-quarter the number in the spermatocyte. 
Within the archoplasm of the spermatid the proacrosomic 
sranules have collected (or according to Meves, Niessing, or 
Moore, now become visible again) (PI. 12, fig. 7, apa); but very 
soon around each proacrosomic granule a clear ring appears, 
so that the granule reposes in a vacuole—the archoplasmic 
vacuole: the proacrosomic granules together with their 
vacuoles in which they lie, now tend to run together, so that 
one obtains the appearance of a number of granules, some 
larger than others (PI. 12, fig. 7, apa). 
At this stage the centrosome is dividing in the cytoplasm, 
near, but outside, the archoplasm (PI. 12, fig. 7, c). 
In the next stage the proacrosomic granules have run 
together so as to form two or three large grains, each surrounded 
by the clear vacuolar ring—the archoplasmic vacuole (PI. 12, 
fig. 8, apc). The whole Golgi apparatus and archoplasm 
gradually passes to the anterior pole of the cell, i.e. that part 
of the cell which gives rise to the head end of the sperm, and 
which most commonly is directed towards the germinal 
epithelium. In PI. 12, fig. 8, the Golgi apparatus and archo- 
plasm are shifting in an upward direction (according to the way 
this cell has been drawn on the Plate). From the posterior 
end of the cell, the axial filament grows out from the centro- 
somes (c! and ©”). 
The next stage in the formation of the acrosome is depicted 
in Pl. 12, fig. 9. A part of the nucleus is shown at n, and the 
Golgi apparatus plus the archoplasm lie nearly in front but to 
one side of the nucleus. The whole apparatus lies in contact 
U2 
