982 J. B.. GATENBY AND J. H. WOODGER 
containing the tail of the sperm became transformed into the 
embryonic part of the germ, the other blastomere into the 
trophoblast. 
Meves (21) likewise suggests something similar for the 
case of Kchinus. That part of the pluteus containing the sperm 
middle-piece is supposed to bud off the Echinus rudiment, 
a very unlikely suggestion indeed. 
Levi (19) in remarking on these facts and suggestions says : 
‘Le ipotesi di Henneguy e di Meves non furono finora 
suffragate da alcun fatto, ed il solo argomento nuovo che io 
adduco, la possibilita della persistenza del pezzo mtermedio 
dello spermatozoo in uno dei blastomeri provenienti dalla 2* 
segmentazione, non contribuisce ancora ad illustrare il si- 
enificato del condrioma maschile nello sviluppo ulteriore.’ 
As one of us pointed out before, the explanation of Henneguy 
for the case of the mammal does not accord with the generally 
accepted interpretation as to the origin of identical twins, 
for if the presence of a middle-piece was a factor of any sort 
of differentiation, the two separating blastomeres would not 
produce the identical twins. 
We see no reason to suppose that the middle-piece Golgi 
apparatus is stripped off the sperm and left outside; there 
seems every justification for the supposition that the apparatus 
is carried into the egg with the mitochondria. What fate hes 
in store for this middle-piece Golgi apparatus is unknown to 
us, nor do the works of van der Stricht, Lams, or Levi bring 
forward any sort of evidence with regard to this point. 
In all probability, the apparatus, like the mitochondria, 
first remains complete and inert and ultimately degenerates, 
after having fulfilled its function, whatever that may be. 
The meaning of the stages in spermatogenesis during which 
most of the mitochondria and part of the Golgi apparatus 
become applied to the middle-piece of the spermatozoon, is 
difficult to understand. If the mitochondria and Golgi 
apparatus of the spermatozoon remain inert, unlike those of 
Ascaris which persist in the egg and live (15), we are forced to 
conclude that the function fulfilled by these bodies is carried 
