CYTOPLASMIC INCLUSIONS OF THH GERM-CELLS 985 
separated from the main Golgi apparatus, and ultimately comes 
to lie in the middle-piece bead referred to in paragraph 2. 
6. The rest of the Golgi apparatus of the ripening sperma- 
tozoon sloughs off. 
7. While all the chromatinic substance of the young 
spermatid eventually goes to form the nucleus of the sperma- 
tozoon, only the majority of the spermatid mitochondria, 
and a very small part of the spermatid Golgi apparatus, form 
the representatives of these cell organs in the ripe spermatozoon. 
8. Attention is drawn to the works of Lams and Doorme, 
van der Stricht, and Levi, where it has been shown that the 
whole middle-piece of the mammalian sperm (Cavia or Vesper- 
tilio) enters the egg at fertilization, but, so far as these authors 
could observe, thereafter remains inert, and is carried whole 
and haphazardly into one of the blastomeres. 
(b) The Formation of the Acrosome. 
9, The acrosome of the spermatozoon of Cavia is formed 
from the proacrosomic granules which are differentiated within 
the archoplasm during the later growth stages of the spermato- 
cyte. 
10. The archoplasm in the spermatocyte of Cavia is covered 
by the Golgi elements or dictyosomes, which in all probability 
are associated with the differentiation within the archoplasm 
of the proacrosomic granules. 
11. Each of the spermatids derived from the spermatocyte 
contain an equal share of Golgi elements, archoplasm, and 
proacrosomic granules. According to Papanicolaou and 
Stockard the latter granules do not disintegrate during mitosis, 
but, keeping their individuality, become scattered in the 
cytoplasm, are subequally sorted out among the daughter 
cells, and eventually come to lie within the re-formed spermatid 
archoplasm. 
12. Hach proacrosomic granule has a liquid-filled space 
formed around it, so that it comes to lie in an archoplasmic 
vacuole. 
13. The several proacrosomic granules within their archo- 
