540 T. A. STEPHENSON 
Endocoelactids. As mentioned on p. 523, the Endocoelactids 
seem to be definitely post-Haleampid and Nynanthean, and 
will here be treated as such. This leaves us with the Gonactini 
idae and Ptychodactidae (see pp. 508, &¢.); and I feel that 
these represent two different side-lines of evolution, not 
necessarily very close together even though both have some 
primitive features, and that in this case it is safer to 
give each a group, the two equal in rank. I therefore 
propose to limit the Protantheae to the Gonactiniidae (in 
the sense taken on p. 505 and exclusive of Boloceroides), 
and to erect a group Ptychodacteae for the Ptychodactids, 
equal in rank to Protantheae and Nynantheae. The Nynan- 
theae I accept as the main tribe, provided it include 
Boloceroides (see p. 506) and the Hndocoelactids (see 
p. 522), and exclude the Edwardsuds and ‘ soft’ corals (see 
p- 510); and provided that not only it, but also the other 
tribes, be re-defined on the sum of their main characters and 
not on the presence or absence of ectodermal muscle in the 
body-wall, simply. 
My suggestion for subdividing Actiniaria is therefore this 
one : 
Tribe 1, PRoTANTHEAE (including Gonactiniidae only, and not Bolo- 
ceroides). 
Tribe 2. PrycHopactTEas (including Ptychodactidae). 
Tribe 3, NYNANTHEAE (including Boloceroides and Endocoelactids, 
and majority-forms, excluding Edwardsiids and skeletonless corals). 
With regard to the other subdivision of Actiniaria into 
Actinunae and Stichodactylmae—I used this, provisionally 
only, in Part I, but am letting it lapse here in favour of the 
above scheme. One feels that these groups have been useful 
as a half-way house, but that in the light of developing know- 
ledge of the group, it is now possible to go farther. The 
‘ Actiniine’ condition is found in all Nynantheae save one 
section; it prevails also in Protantheae, Ptychodacteae, and 
most corals. ‘ Stichodactylinism’ occurs in Ceriantharia and 
a few corals, and in one set of Nynantheae. There are, however, 
among Nynantheae, four quite distinct sets, seemingly repre- 
