1023 
To estimate this relative development Artnur KeitH*) has intro- 
duced the comparison of the capacity with the “palatal area’, this 
area being the space bounded by the outer margins of the crowns 
of the teeth in the upper jaw and a line joining the posterior margin 
of the upper third molar teeth. He found this area of the upper 
dental arcade for a female chimpanzee skull, of a capacity of 320 
em.*, equal to 36.5 cm.*; hence to 1 cm.’ of palatal area came 
8.7 cm.* of brain capacity. The upper palatal area of a Tasmanian 
skull was 36.8 cm.’, the capacity of this skull was 1350 cm.’, 
which gives a ratio of 1:36.7. For the Homo neandertalensis of 
Gibraltar*) Keira found for these values 31.6 cm.* and 1200 em.’, 
and the ratio 1:38, but for the Aurignac-man of Combe-Capelle the 
ratio is 1:53, about that of modern Englishmen, viz. 1 : 56.3, 
with 26.6 cm.” palatal area and 1500 cm.’ capacity. 
With pretty great accuracy — as only the crowns of the incisors 
and the crown of the right m, fail — the palatal area of the 
Wadjak-man II may be determined at 41.4cm.*. That of Wadjak I, 
in which only few tooth-crowns have been left, measures about 
35 cm”. Through its relatively small size this palate presents a 
striking difference from that of Wadjak II, which is one of the 
characters that lead me to assume that the first found fossil remains 
belonged to a woman, the second to aman. Other female characters 
of Wadjak I are: the more reduced form of the teeth (the upper 
m, and m, are almost perfectly three-cusped), the smaller dimensions 
of the comparable parts of the skull, though not in the same degree 
smaller as the palate, the less pronounced superciliary ridges and 
the forehead that does not recede so much, the orbits which are 
higher with respect to their breadth, the somewhat slighter develop- 
ment of the muscle attachments, the more rounded form of the 
occiput, the somewhat slighter lophocephaly and dolichocephaly, in 
so far as the latter can be judged from the fragments of the second 
skull. 
If for the fossil woman of Wadjak 44.3 em.” brain capacity comes 
to 1 cm. of palatal area, it may be assumed that for the man, who 
had a much larger palate, but probably also a larger neurocranium, 
this ratio was smaller. Putting his cranial capacity 100 em.’ higher 
1) ARTHUR Keir, The Antiquity of Man. (London 1920), p. 97, 151, 328. 
2) For the Homo neandertalensis of La Chapelle-aux-Saints I calculate an upper 
palatal area of 38 cm? after the reconstructive drawing of Bouvre, which with 
1626 cm’ Broca- or 1530 real cranial capacity yields the ratio 1:40.38. But the 
normal palatal area may have been somewhat larger than that of this man, who 
had early lost his teeth for the greater part. 
