BY C. D. GILLIES AND C. T. WHITE. 43 
due to the fact that the material was not immediately 
preserved in the formalin after collecting, for the specimens 
were not originally gathered for sectioning. 
With reference to the hypogynous gland, Longman 
and White (1918, p. 162), state that ‘‘in practically every 
flower the hypogynous gland is divided into four or five 
segments (usually five) and two of these are much elongated 
into supplementary style-like processes,” and later (p. 164), 
* There is no evidence of a graduated change from the 
tiny segments of the hypogynous gland, and it is therefore 
thought that this marked modification is better expressed 
as a mutation than as a variation.” After a _ careful 
examination of our sections; we find that we cannot support 
the opinion that these style-like processes have any 
connection with the hypogynous gland, as it appears to us 
most conclusively on morphological and histological grounds, 
that they are neither hypogynous gland-segments nor 
a mutation from them. On the contrary, their resemblance 
to the style is very pronounced in regard to the following 
important features, viz. (a) general shape, (b) stigmoid 
extremity, and (c) microscopic structure. In Fig. 1. it 
will be noticed with reference to the histology of the stipes 
{which is similar to that of the style) and the style-like 
processes, that these organs are chiefly composed of lightly 
staining parenchyma (par.) surrounding a delicate central 
strand of vascular tissue (v.b.), and containing a few 
scattered deeply-stained cells (c). Contrasting and alter- 
nating with the stipes and the style-like processes are the 
segments of the hypogynous gland (hl, h2; h3), which stain 
deeply and consequently are conspicuous structures in 
section. We therefore conclude that the elongated organs 
referred to by Longman and White as segments of the 
hypogynous gland are really aborted styles and that their 
development has caused splitting of the hypogynous gland 
into distinct segments. 
Early this year, one of us obtained typical flowers of 
Buckinghamia celsissima from the Brisbane Botanic 
Gardens, and sections for comparison were made. It will 
be observed that the hypogynous gland (Fig. 2, h) is entire 
and semi-annular in shape and that aborted styles are not 
present. 
