( 900 ) 
there sends out a series of fibres to the Ili nucleus (compare also 
EpiNGER and WALLENBERG ')). 
In connection with the more frontal migration of the dorsal nucleus 
which was found in all Teleosts, | may recall here the fact that 
DE LANGE found the decussation of the dorsal tecto-bulbar fibres also 
more frontal than the decussation of the ventral set. 
Regarding the III root I will mention that a great number of 
decussating fibres originate from the posterior part of the IIT nucleus 
bordering on the trochlear nucleus, which is interesting in connection 
with the fact that the IV fibres have also a decussating character. 
The trochlear nucleus shows us a further stadium in the process 
already indicated in the shark, viz. the frontal shifting of its cells 
which here lie entirely in front of their root-entrance. Although this 
may partly result from a backward displacement of the root in some 
Teleosts, a more frontal shifting of the IV cells is also very probable, 
as is seen from the fact that the IV nucleus in Lophius has also a 
more frontal position in regard to the [IL root and V root and there- 
fore, with regard to these points also, the shifting of the IV nucleus 
can be affirmed, equally in all Teleosts. (Figs. 3 and 11). 
The position, which the IV nucleus occupies with respect to its 
root-entrance in the bony fishes is strongly suggestive of that in human 
beings (cf. _MarBure ©) where the nucleus also lies entirely frontad 
thereof. It has been found by van VALKENBURG *) that this secondarily 
produced relation is sometimes shown by a cauda/ vemnant of the 
IV nucleus, which he designates “nucl. LV posterior” (cf. JACOBSOHN*). 
With regard to the trochlear-root the following point may be 
mentioned. 
Earlier investigators had already observed that the trochlear-root 
traverses the brain-stem with 2 roots (Salmo, Haier *) Gadus, Kar- 
PERS *)) in some bony-fishes in contrast to other bony-fishes (e. g. 
1) Vorlesungen 7te Auflage. — Beiträge zur Kenntnis des Gehirns der Telecstier 
und Selachier. Anat. Anzeiger, Vol. 31, P. 369. 
2) MarsurG, Mikroskopisch-topographischer Atlas des menschlichen Zentralnerven- 
systems. 
3) C. T. van VALKENBURG: These Proc. June 25, 1910. 
4) Jacopsonn, Verhandl. Preuss. Akad. 1909. 
5) HALLER. Vom Bau des Wirbeltier-gehirnes. Morphologisches Jahrbuch Bnd. 
26, 1898, p. 508. | cannot affirm however HALtrr’s statement that a part of the 
IV root originates from the Purkinje-cells of the cerebellum. Nor did [ see a 
“kontinuierliche Zusammenhang dieses Kernes (LV) mit dem rostralwärtigen Ende 
des oberen motorischen Trigeminuskernes” (l.c. p. 505). 
6) Karpers. The structure of the Teleostean and Selachian brain. Journal of 
Comparative Neurology. Vol. XVI, 1906, p. 62. 
