in which I found the 4 toe inside this field, there were always 
variations in the plexus extant pointing to the fact that the extremity 
was designed postfixally. The foremost boundaryline of this field first 
coincided dorsally and ventrally from the medianlines for some 
distance with the posterior boundary of the 4" lumbarsegment. I 
found likewise in different cases here an overlap of about '/, em- 
These will certainly be the experiments that succeeded best. Cari- 
‘ature formation is, after all, likewise with segments of strychnine by 
no means rare. 
Sacralis 2 can also send a smaller or larger slip to the upperleg. 
A single word about the axile lines of SHeRRINGTON called by Bork 
differentiation-boundaries. 
We know for certain that originally the 4 lumbarsegment and 
the 1st sacralsegment, have not verged to each other. The experiment 
taught me however, that the overlap is much stronger than was 
originally supposed, but the mutual overlap of succeeding segments is 
not so strong, that the 4 lumbarsegment and the 1' sacralsegment, 
between which 3 segments are lying, can originally have verged on 
each other. 
With the thoracic lumbarsegments | found in the dorsal region 
overlap of every two segments that were separated in succession by 
another, and in the ventral region there was still overlap of 2 
segments that were separated in succession by 2 other segments. 
Nowhere did I see 2 segments verge on each other that were separated 
in succession by three other segments. We know consequently for 
certain, that the 4 lumbarsegment and the {st sacralsegment origin- 
ally have not overlapped each other and have not verged on each 
other, but that there has existed between the caudal boundary of 
the 4 lumbarsegment and the cranial boundary of the 1st sacral- 
segment a zone of a certain width. After the development of the 
extremity these 2 boundaries of dermatomes have approached each 
other, and the two dermatomes between which there was originally 
a distance, have even for a narrow zone overlapped each other. 
This fact was already known from SHERRINGTON’s experiments. SHER- 
RINGTON points to the fact that the overlap at the axile lines has 
much resemblance to the dorsal and the ventral ‘crossed-overlap”. 
About as far as the boundary between the inferior and the central part 
of the upperleg these two dermatome-boundaries coincide ; afterwards 
the anterior boundary of sacralis 1 assumes a more slanting posterior 
direction. The superior part of the 5‘ lumbar dermatome reaches nearly 
this height, so that then the length of the axial line corresponds with 
the distance, along which these two dermatome boundaries coincide, 
