Occasional Papers of the Museum of Zoology 17 



QuADRUivA NODULATA Rafinesque, 1820 

 Type locality: Kentucky River. 



Obliquaria (Quadnila) nodulata, Rafinesque, '20, p. 307, pi. 81, f. 

 17-18. 



Unio pustulatus Lea, '31, p. 79, pi. 7, f. 9. 



U. nodulatus Raf. = U. pustulatus Lea, Conrad, '34, p. 70; '37, p. 80. 



Quadrula pustuhta (Lea), Simpson, '14, p. 856. 



Quadrula nodulata (Raf.)=(3. pustulata (Lea), Vanatta, '15, p. 

 557 ("type" examined). 



Quadrula nodulata (Raf.), Utterback, '16, p. 57. 



Obli. nodulata Raf. = Q. pustulata (Lea), Walker, '18^', p. 168 ("if 

 identifiable"). 



The original description of nodulata mentions and the 

 (poor) figure shows the characteristic posterior '"wing" of 

 this shell; also the arrangement of the distant (text) tubercles 

 in two rows is suggested in the figure. A nodulous shell with 

 these characters can only be the pustulata of Lea. Thus, 

 Rafinesque's species is recognizable, which is confirmed by 

 Vanatta's examination of the so-called Rafinesque-Poulson 

 "type." 



Quadrula mETanevra Rafinesque, 1820 



Type locality: Kentucky River. 



Obliquaria (Quadrula) nictaiievra Rafinesque, '20, p. 305, pi. 8r, 

 f. 15-16. 



Unio nodosus Barnes. '23, p. 124, pi. 6. 



U. metanevra Raf. = [/. tiodosus Bar., Conrad, '34, p. 70; Ferussac, 

 '35, P- 27; Conrad, '35, p. 10, pi. 5, f. 2. 



Theliderma metanevra (Raf.), Swainson, '40, p. 378. 



Quadrula metanevra (Raf.), Agassiz, '52, p. 48; Simpson, 14, p. 834. 



There has never been any doubt, since the time of Conrad, 

 as to the identity of this species, and indeed Rafinesque's 

 description and figure are identifiable. 



However, this species should not be regarded as the type of 

 the genus Quadrula. It belongs in the group of U. cylindricus 

 Say, for which Orthonymns Agassiz ("52) is to be used as 

 sectional (or subgeneric) name (see above). 



