GOLOI BODIES OF A COCCIDIAN 387 



takes no part in the process. We have an apparatus in the 

 coccidian which acts normally at dictyokinesis. It is evident 

 that in this protozoon we have the typical apparatus already 

 formed and established. 



The search for a primitive apparatus must be carried out 

 among forms other than the Sporozoa, as Hirschler's and our 

 work amply shows. 



The Golgi bodies probably arose in connexion with the ter- 

 minal bead of the flagellum of some primitive flagellate. The 

 outer layer of the bead might have been differentiated to form 

 a lipoid store-house or elaborator of the energy-yielding 

 materials necessary for the nutrition of the locomotor organ. 

 From its primitive position in the metazoan cell, always 

 associated at some time with the centrosome-centrosphere com- 

 plex, we cannot but believe that in the early history of the 

 cell the Golgi apparatus and the centrosome were evolved side 

 by side, or thea pparatus from the centrosphere, in some way. 

 This speculation can only be tested when further evidence is 

 produced. An important field, altogether neglected hitherto, is 

 opened up to protozoologists. The latter more than any of 

 their fellow biologists are interested in the architecture of the 

 organisms which they study, and they should attack the 

 problem with the special methods explained elsewhere (5). 



Summary. 



1. There is a true Golgi apparatus in the Coccidia (PI. 21, 

 figs. 11, 12, 16, 17, 18, 21). 



2. It consists of separate dictyosomes or crescentic rods, 

 with the power of fission as in metazoa (PI. 21, figs, 2, 12, 17). 



3. During growth the excentric Golgi apparatus (PI. 21, 

 fig. 5) becomes larger and tends to spread out in the cell (PI. 21, 

 figs. 12, 13). 



4. During division of the schizont the Golgi elements are 

 attracted into subequal groups of dictyosomes and granules 

 around each nucleus, as happens in most metazoan cell- 

 visions. 



