298 C. CLIFFORD DOBELL. 
masses, whose number varies according to the size of the 
animal, and which multiply by a simple division (text-fig. 16). 
They cannot properly be called nuclei. They should be 
regarded, I think, as composing a nuclear apparatus consist- 
ing of scattered fragments of chromatin—a chromidial system 
—as in some bacteria (e. g. B. flexilis, Dobell, ’08a). Inlarge 
animals they are present in immense numbers, but at no 
period do they—individually—possess the attributes of a 
formed nucleus. 
In some Protozoa nuclear reduction by chromidia formation 
takes place in a gamete preparatory to conjugation (e.g. 
macrogametocyte of Adelea (Siedlecki, ’99), and in 
TEXxT-FIG. 16. 
Large Siedleckia nematoides (from Aricia fctida). 
C. chromatin fragments in the cytoplasm. (Original.) 
Monas (Prowazek, 703). ‘Their meaning is bound up with 
the general problem of nuclear reduction, and I shall say no 
more about it here. 
(B) CHRomripIA IN BAcTERIA. 
In spite of the great discussion which has raged—and still 
rages—round the problem of the bacterial nucleus, there is a 
large and growing body of evidence to show that some, at 
least, of the granular inclusions in bacteria consist of chro- 
matin (cf. Guilliermond, ’07). In part, however, the granules 
(““metachromic granules,” “red granules,” “volutine granules,” 
etc.) probably consist of some reserve material (cf. Guillier- 
mond, ’06, 707). It can hence be said that certain bacteria 1 
1 And probably also Cyanophyceex, 
