536 GEOFFREY SMITH. 
of the stomach by a wide common opening. The histological 
character of these tubes varies greatly according to the 
condition of metabolism. A transverse section through the 
middle of a tube, when the animal is starved, shows a regular 
lining of tall columnar cells, the majority of which have 
hyaline, reticular cytoplasm, staining pink with eosin, and 
granular nuclei (Pl. 12, fig. 8). These cells are mainly 
absorptive in function. Scattered among these cells are 
narrower cells with rather coarsely granular cytoplasm, which 
is darkly coloured with hematoxylin. ‘These cells are 
special gland-cells and probably secrete a ferment. 
At the ends of the tubes the nuclei are greatly crowded, 
but both kinds of cells can be recognised. 
After feeding heavily, the histology of the tubes changes 
(Pl. 12, fig. 9). The special gland-cells are no longer recog- 
nisable, and the absorptive cells lose their definite cell-outlines 
and are distended with oily globules. Certain of the cells 
consist merely of an envelope containing an immense vacuole 
with a darkly staining nucleus flattened on one side. The 
liver of Anaspides therefore has, at least, two distinct 
functions ; it produces digestive ferments which are poured 
into the stomach, and it also plays an important part in the 
absorption and storing of assimilated material. 
If we compare the alimentary tract of Anaspides with 
that of other Malacostraca we see that the structure of the 
stomach points rather to Kucaridan affinities. The presence 
of dorsal diverticula at the juncture of endodermal mid-gut 
with stomodzeum and proctodeeum is a Decapodan character, 
since in the ‘“‘Schizopoda,” i.e. Kuphausiide and Myside, 
etc., there is never a diverticulum between mid-gut and 
proctodeeum. We have seen that there is also another diverti- 
culum in the middle of the mid-gut, and this character is, as 
far as we know, peculiar to Anaspides and its immediate 
allies. 
The liver, although in certain respects peculiar, is nearer 
to the Eucaridan plan than to the Peracaridan, since in the latter 
there is typically present a glandular ridge upon which the 
