OBSERVATIONS ON TKYl'ANOPLAS:\rA CONGKRf. 493 



seineu einzelneu Phasen im Leben niclit verfolgeu konueu. 

 Wie icli sclion oben angegeben habe^ trifffc mnn relativ selten 

 sicb verinebrende Individueu an. Vorzugsweise siud es 

 Tierc, bei deiien die Teilung des clironiatisclien Apparates 

 und des Plasmas^ sowie die Bilduiig der lokoixiotoi-isclien 

 Organellen bereits beendet siud und die uiir iiocli uiit ihrer 

 liintei"eu Eudeii zusauimenhangeu, eiue Pliase^ die zeitlich 

 Ijingste im Laufe der Teilung- zu seiu sclieint.'' It is parti- 

 cularly over this last point, liowever, that a. great de:il of 

 caution should be exercised. In well-infected smears it is 

 an exceedingly common occurrence to find two trypanoplasma 

 lying iu a position which suggests division, but unless there 

 is some absolutely distinctive feature, e.g. as regards the 

 structure of the nuclei, which can be definitely connected 

 with a corresponding structure in an undoubted dividing 

 form, I feel that it is always most hazardous to interpret 

 these appearances as division st.iges. On the other hand, 

 the differences between the division of T ry pan oplasma 

 helicis, as described by Friedrich, and that of Trypano- 

 plasma congeri, seem to be of an absolutely fundamental 

 charactei-. In the first place the karyosome, which is so 

 characteristic a feature of the trophonucleus of most trypano- 

 plasmas, is entirely absent iu Ti-y panoplasma helicis, and 

 in correlation with this fact the division of the trophonucleus 

 appears to consist in a simple constriction of the large 

 vacuolar trophonucleus with its scattered chromatin granules 

 (p. 387), The division of the kinetonuclens is said to be 

 longitudinal (p. 385), but the figures of this process seem 

 hardly convincing. The behaviour of the flagella, again, 

 seems to be very complicated, since it is said on p. 390: 

 " Nachem die fiir die neue Zelle notwendigen Teile entvvickelt 

 sind oder der Anlage nach vorhanden siud, riicken die Kerne 

 und Blepharoplasteu auseinander." " Dasselbe geschicht 

 mit den Geisselursprungsstellen, die alsdann im die Niihe 

 des Blepharoplasteu verlagert werden. Dabei bildet sich die 

 der alten undulierenden Membran zunachst gelegene Geissel- 

 anlage sur vordereu Geissel eines neues Tieres aus, wahrend 



