674 H. M. WCODCOCK. 



it is cleai-j I think, that this structm-e has really nothing to 

 do with a tlagelluin. The line is usually most prominent 

 in parasites which show numerous retl-staiuing (probably 

 chromatoid) granules in the cytoplasm; and, in suitable 

 instances, it can be seen quite well that it is situated at the 

 edge of the endoplasmic intrusion in the membrane (figs. 

 42, 47,49). Further, when present, it can usually be traced 

 right along the course of the membrane from end to end. 



If we had to deal here with a case of division or formation 

 of a new flagellum, individuals showing either an earlier or 

 later phase in the process might be expected to occur, for 

 this appearance is not at all infrequent; but I have not 

 found any such. Again, in most cases, there is not the 

 least indication of nuclear division. Lastly, in one of the 

 exceptionally few instances where any indications of division 

 are present, in addition to the kinetonucleus having divided 

 into two, the true tlagellum can be seen to be itself double 

 for a shoi't distance near its proximal end, probably as a 

 result of splitting (tig. 54) .^ The granular chain is also 

 present, and, as before, quite separate from the flagellum. 

 Hence there is no reason for regarding this structure as in 

 any J way connected with a flagellum, much as it simulates 

 one at times. 



The small stumpy tryiDanosome in fig, 41 shows what is 

 probably an early stage in the development of this line. 

 Here there is a row of red-staining granules, quite separate, 

 and not closely apposed to constitute a chain, which run 

 parallel to the flagellar border, doubtless at the limit of the 

 endoplasm. The granules are apparently quite similar to 

 others which are seen in the general cytoplasm. I have no 

 idea what is the explanation of this aggregation of chroma- 

 toid granules into a compact chain, lying in the position 

 described. I have never seen it either in the ordinary 

 definitive trypanosomes or in parasites of the other large type. 

 I may add that I have observed the same feature in the case 

 of a trypanosome from a blackbird (Turd us merula), at 

 ' C f . also the micro-pliotograph reproduced in fig. D. 



