9 



liaiid and tlie contraclile iihrils. the sarcostyles on the other hand, 

 cüukl he stated with accuracy. I often j^ot tlio impression that tlie 

 delicate nerve-tibrils of tiie motor nerve-ending, after entering the 

 mnscle-tibre, run round tiie sarcoplasmatic prominence, encircHng it, 

 and then follow the direction of the sarcostyles for some distance 

 before breaking up in endrings or endnets. These motor nei've-endrings 

 1 found both in the circular tibres of the sphincter pupillae and in 

 the radiating fibres. The innervation of the membrane of Brich I 

 could not make out. 



Beside these motor nerve-endings at the end of medullated nerve-fibres 

 tliere exist in the sphincter pupilke and in the musculus ciliaris of 

 the bird's eye just as in the voluntary sceletal muscles the very 

 delicate, non-medullated nerve-fibres with their small, loosely arranged, 

 delicate endings, entering the muscle-fibers independently of the 

 motor nerve-terminations mentioned above, which I described in the 

 voluntary muscles of the body as "accessory nerve-terminations", 

 and which could be traced to the nervus sympathicus. In fig. 8, 9, 

 and JO are given some examples of these accessory nerve endings 

 on muscle-fibres of the sphincter pupillae and the musculus ciliaris. 



As far as I could gather from my preparations, the delicate non- 

 medullated nerve-fibres ending in the small "accessory" terminations 

 on the muscle-fibres, remained independent of the medullated nerves. 



Whether we are entitled to ascribe to these accessory nerve-fibres 

 even here an influence on the tonus of the iris- and ciliary muscles, 

 and what are the relations of these nerve-fibers to the sympathetic 

 nerve, are cpiesfions not to be debated here. There is here a wide 

 field lying open for experimental study. 



In conclusion some words may be added about the question, 

 whether ganglioiicells are [)resciit in the corpus ciliare or in the 

 stroma of the iris or not. Several authors (e.g. Retzius) denied the 

 existence of these ganglioncells, others, as C. Krause, H. Muller, and 

 in later years especially .Geberg, described small groups of ganglion- 

 cells and separate cells appearing alongside the nerve-fibre bundles, 

 others, as Agababow, found ganglioncells only in the course of the 

 vasomotor nerves. Finally Inglis Pollock found in J9J2 that after 

 the exstirpation of the ganglion ciliare or of the superior cervical 

 sympathetic ganglion the nerves of the corpus ciliare and iris did 

 not degenerate. This fact woidd point to the conclusion, that in the 

 corpus ciliare and the iris the ganglioncells are as al)undanf as they 

 are in the terminal sympathetic ganglia in the intestinal wall in the 

 nerve plexus of Auerbach and Mkissnek. 



In my preparations of the corpus ciliare and iris, stained after 



